aL e; (AG | | | in CAT. AUG 1 2 2004 LIBRARIES a at Volume 78 (1) 1999 = Supplemento al Bollettino della Societa Entomologica Italiana, 131 (3) (30.10.1999) 30 aprile 2000 SOCIETÀ ENTOMOLOGICA ITALIANA Sede in Genova, via Brigata Liguria, 9 presso il Museo Civico di Storia Naturale M Consiglio DIRETTIVO 2000-2001 Presidente: Augusto Vigna Taglianti Vice Presidente: Mario E. Franciscolo Segretario: | Roberto Poggi Amministratore: Giovanni Dellacasa Direttore delle Pubblicazioni: Riccardo Sciaky Consiglieri: Baccio Baccetti, Sebastiano Barbagallo, Claudio Canepari, Attilio Carapezza Achille Casale, Fabio Cassola, Mauro Daccordi, Giulio Gardini Giuseppe Osella, Fernando Pederzani Enrico Ratti, Stefano Zoia Revisori dei Conti: Enzo Bernabò, Enrico Gallo, Giuliano Lo Pinto, Revisori dei Conti supplenti: Ducezio Grasso, Sergio Riese Bibliotecario: Giulio Gardini Comitato di redazione: Achille Casale, Fabio Cassola, Mauro Daccordi, Mario E. Franciscolo, Roberto Poggi, Riccardo Sciaky, Augusto Vigna Taglianti, Stefano Zoia Segreteria di Redazione: Stefano Zoia M CONSULENTI EDITORIALI Nits M@LLER ANDERSEN (K@benhavn) - PAOLO A. AUDISIO (Roma) - GEORGE E. BALL (Edmonton) - EMILIO BALLETTO (Torino) - MARCO A. BOLOGNA (Roma) - BARRY BOLTON (London) - PIETRO BRANDMAYR (Cosenza) - MARIO COLUZZI (Roma) - ROMANO DALLAI (Siena) - THIERRY DEUVE (Paris) - ALESSANDRO FOCARILE (Medeglia) - ERNST Heiss (Innsbruck) - MANFRED JACH (Wien) - MARCELLO LA GRECA (Catania) - VOLKER MAHNERT (Genève) - Luigi MASUTTI (Padova) - ALESSANDRO MINELLI (Padova) - CLAS M. NAUMANN (Bonn) - Lazio Papp (Budapest) - SANDRO RUFFO (Verona) - VALERIO SBORDONI (Roma) - KONRAD THALER (Innsbruck) - STEFANO TURILLAZZI (Firenze) - S. BRADLEIGH VINSON (College Station) - JEFF F. WAAGE (Ascot) - ADRIANO ZANETTI (Verona) - ALBERTO ZILLI (Roma) - PETER Zwick (Schlitz). ISSN 0037 - 8747 Fondata nel 1869 - Eretta a Ente Morale con R. Decreto 28 Maggio 1936 Volume 78 (1) 1999 30 aprile 2000 REGISTRATO PRESSO IL TRIBUNALE DI GENOVA AL N. 76 (4 LUGLIO 1949) Prof. Cesare Conci - Direttore Responsabile Spedizione in Abbonamento Postale 50% - Quadrimestrale Stampato da PolyGrafika, Via Ciro Menotti 11/D, 20129 Milano SOCIETÀ ENTOMOLOGICA ITALIANA via Brigata Liguria 9 Genova iy ‘gr 4 h Paci Ranco we gel, vy de | | tria © perso ie iit we HET x | — Li Ge nr th FIL: ap et) a . PU i +a Da CA ; Ws oy ai UE We n dia they winded ~~ a È WE oy i dis à APE at) s ib CRC RUE rm —— pos 5 i ae 1 mi tai man LU CORTE : Ae 2 È a wy i 7 y hi LIU Er Pi any i fl ie or Wks tilt i mi UT die A ET I | i 2 AGRO f ini ll (LI na tr Dali dia 1 mi | VA ha er ir A ta y i fil it! VE ei i ri an Sa ALLEA MALE CT: VUS h 1 fin diay È (D } : sf Tos 1 Tie Ms "i MEET pi ni sci À i LL là “ ra a Mn hi fi j En I Ne Y ee if ” Ne oo Ln A è À l ni) i F x IM hi (x be avis yy l WP = Li L: Si | 7 A “ti épi) À Au a | MYAR) = Gaal à * ic 1 Pas ry il Zen Ha de cheney a Eames | La | met tote 144 SU tar NU CN 4 br LUCE Vor via [METTE 46 ani vok - bia a bi iui M ne My ge Dr i LA. pia in Mica; gh "I ie MBA ARENA UT è 00 Huhn We cal Ma! i i nr Ù reste pese Tr un state dre Mem. Soc. entom. ital., 78 (1): 3-4. 30 aprile 2000 Peter John MAZZOGLIO Felice Musso (27.11.1914 - 13.VII.1999) Riassunto - Viene qui ricordata la figura di Felice Musso, entomologo dilettante e primo dipen- dente dell’Istituto di Entomologia agraria dell’ Università degli Studi di Torino. Abstract - Felice Musso (27.11.1914-13.V11.1999). The figure of Felice Musso, an amateur entomologist and the first employee of the Institute of agri- cultural Entomology of the University of Turin, is here recalled. Entomologo e apicoltore dilettante e filatelico; la ricca collezione di francobolli a soggetto entomologico, andata purtroppo perduta, rappresentò il suo unico contributo pub- blico in occasione del Simposio Internazionale dell’ Apimondia su “Flora mellifera e impol- linazione”, tenutosi a Torino nel 1972!, ripetuto nel Convegno su “Passato e Presente dell’ Apicoltura Subalpina” tenutosi anch’esso a Torino dieci anni dopo. Per l’entomologia torinese, con Felice Musso scompare anche il primo dipendente I Vidano C., 1977 - Api e apicoltura nella filatelia internazionale - L’apicoltore moderno, 68(1): 15-19. 4 MAZZOGLIO dell’Istituto di Entomologia agraria, fondato nel 1936 da Athos Goidanich. Infatti, Felice vi si trasferì dalla Clinica Medica Universitaria dell'Ospedale Molinette nel 1938 e, per molti anni, fu l’unico aiuto del direttore in campo e in Istituto. Alternò l’impegno militare con il lavoro entomologico all’Istituto, partecipando alle ricerche sull’entomofauna del riso, della vite, sui cinipidi, sugli scolitidi e su vari insetti dannosi d’attualita. Con lo scoppio della guerra fu impegnato come marconista a Torino, in Valle di Susa, in Francia e Liguria, quindi nella batteria contraerea a Leinì. Con l’armistizio entrò nella 10° Brigata d’assalto Garibaldi “Piacibello” a Casalino di Mombello Monferrato in Val Cerrina. Anche in quest'altra veste servi la causa come staffetta per portare ordini in tutto il Piemonte. Alla fine della guerra rientrò all’ Università, aderì al movimento sindacale e fu fra i fondatori della CGIL dell’ Università di Torino. Questo periodo, più tormentato, lo portò dalla Facoltà di Agraria a quella di Farmacia. Il ritorno all’entomologia si ebbe nel 1970. Richiamato da Carlo Vidano, direttore dell’appena fondato Istituto di Apicoltura, lavorò con passione con le api fino al 1976, quan- do dovette essere pensionato per limiti di età, ma la sua presenza all’Istituto fu continua, anche in seguito. Sempre servizievole, non mancava di trascorrere giornate intere, svol- gendo commissioni e lavoretti fino a pochi anni fa, quando la cecità lo privava del bene più prezioso. | Fu membro della Società Entomologica Italiana dal 1971 al 1998. Indirizzo dell’Autore: P. J. Mazzoglio, Di.Va.P.R.A. Entomologia e Zoologia applicate all’ Ambiente “Carlo Vidano”, via Leonardo da Vinci 44, I-10095 Grugliasco TO, Italia. Mem. Soc. entom. ital., 78 (1): 5-70. 30 aprile 2000 Luca TOLEDANO Systematic notes on the palaearctic Bembidiini with particular reference to the fauna of China (Coleoptera Carabidae) Abstract - The systematics of species of the eastern palaearctic subgenera Philochtemphanes Netolitzky, 1942, and Aureoplataphus Netolitzky, 1942, is here reviewed. The subgenus Josefia n. subg., with five new species, and the subgenus Bembidromus n. subg., with one new species, are here described. Some taxa belonging to the subgenera Ocydromus s.l. and Bembidion are here descri- bed as new. Many Bembidion species, mainly from the eastern palaearctic region, are discussed and arranged into known subgenera, in an attempt to reorganize in a modern way the great number of species previously not included in any subgenus. Brief discussions on the significance of some diagnostic characters for groups within the large subgenus Ocydromus s.l. are also given. Some keys for the determination of subgenera, species groups and species, and some synonymic notes are also given. The nomenclatorial acts proposed in this work are: Taxa described as new (in systematic order): B. (Philochtemphanes) brancuccii n. sp. (China); B. (Philochtemphanes) daliangi n. sp. (China); B. (Bembidion) humerale petribulirschi n. ssp. (Turkey); B. (Josefia) n. subg. (China); B. (Josefia) belousovi n. sp. (China); B. (Josefia) luisae n. sp. (China); B. (Josefia) pieroi n. sp. (China); B. (Josefia) taguense n. sp. (China); B. (Josefia) shugela n. sp. (China); B. (Microsinocys) schuelkei n. sp. (China); B. (Bembidromus) n. subg. (China); B. (Bembidromus) panda n. sp. (China); B. (Aureoplataphus) jaechi n. sp. (China); B. (Ocydromus) schoenmanni n. sp. (China); B. (Ocydromus) radians puetzi n. ssp. (China); B. (Ocydromus) radians shaanxianum n. ssp. (China); B. (Ocydromus) moritai n. sp. (China); B. (Ocydromus) baehri n. sp. (China); B. (Ocydromus) echarouxi n. sp. (China); B. (Ocydromus) maddisoni n. sp. (China); B. (Ocydromus) roberti n. sp. (China); B. (Ocydromus) muellermotzfeldi n. sp. (China). Species transferred here to Bembidion subg. Philochtemphanes pera 1942: B. (Emphanes) perditum Netolitzky, 1920. Species attributed here to Bembidion subg. Ocydromus Clairville, 1806: B. radians Andrewes, 1922; B. charon Andrewes, 1926. Species attributed here to Bembidion subg. Peryphophila, Netolitzky, 1942: B. endymion Andrewes, 1935. New synonymies: B. tienmushaniense Kirschenhofer, 1984 = B. perditum Netolitzky, 1920 n.syn.; B. nanpingense Kirschenhofer, 1984 = B. peleum Jedlicka, 1933 n.syn.; B. niedli Jedlicka, 1965 = B. (Ocydromus) yunnanum Andrewes, 1923 n.syn.; B. kinfushanum Jedlicka, 1958 = B. (Ocydromus) hasurada Andrewes, 1924 n.syn.; B. (Pseudolimnaeum ) jedlickanum Toledano, 1999 = B. (Pseudolimnaeum) arnosti Nakane, 1978 n.syn. Redescriptions: B. (Philochtemphanes) Netolitzky, 1942; B. (Philochtemphanes) exquisitum Andrewes, 1923; B. (Philochtemphanes) hansi Jedlicka, 1932; B. (Philochtemphanes) perditum Netolitzky, 1920; B. (Aureoplataphus) Netolitzky, 1942. B. (Aureoplataphus) aureofuscum Bates, 1883. Riassunto - Note sistematiche sui Bembidiini paleartici con particolare riferimento alla fauna di Cina (Coleoptera Carabidae) In questo lavoro vengono sottoposti a revisione i sottogeneri Philochtemphanes Netolitzky, 1942 6 TOLEDANO ed Aureoplataphus Netolitzky, 1942 della regione paleartica orientale. Sono qui descritti il sotto- genere Josefia n. subg. insieme con le sue cinque nuove specie, e il sottogenere Bembidromus n. subg. insieme con la sua unica specie. Vengono descritti qui anche altri nuovi taxa appartenenti ai sottogeneri Ocydromus e Bembidion s.str. In questo lavoro vengono discusse molte specie di Bembidion principalmente della regione palear- tica orientale inquadrandole in sottogeneri noti, nel tentativo di riorganizzare in un modo moderno la grande quantità di specie attualmente non attribuite ad alcun sottogenere; anche il grande sotto- genere Ocydromus è trattato in questo lavoro con brevi discussioni sul significato di alcuni carat- teri diagnostici. Sono anche fornite chiavi per la determinazione di sottogeneri e specie, ed alcune note sinonimi- che. Gli atti nomenclatoriali introdotti in questo lavoro sono: Nuovi taxa descritti (in ordine sistematico): B. (Philochtemphanes) brancuccii n. sp. (Cina); B. (Philochtemphanes) daliangi n. sp. (Cina); B. (Bembidion) humerale petribulirschi n. ssp. (Turkey); B. (Josefia) n. subg. (Cina); B. (Josefia) belousovi n. sp. (Cina); B. (Josefia) luisae n. sp. (Cina); B. (Josefia) pieroi n. sp. (Cina); B. (Josefia) taguense n. sp. (Cina); B. (Josefia) shugela n. sp. (Cina); B. (Microsinocys) schuelkei n. sp. (Cina); B. (Bembidromus) n. subg. (Cina); B. (Bembidromus) panda n. sp. (Cina); B. (Aureoplataphus) jaechi n. sp. (Cina); B. (Ocydromus) schoenmanni n. sp. (Cina); B. (Ocydromus) radians puetzi n. ssp. (Cina); B. (Ocydromus) radians shaanxianum n. ssp. (Cina); B. (Ocydromus) moritai n. sp. (Cina); B. (Ocydromus) baehri n. sp. (Cina); B. (Ocydromus) echarouxi n. sp. (Cina); B. (Ocydromus) maddisoni n. sp. (Cina); B. (Ocydromus) roberti n. sp. (Cina); B. (Ocydromus) muellermotzfeldi n. sp. (Cina). Specie trasferite qui a Bembidion subg. Philochtemphanes Netolitzky, 1942: B. (Emphanes) perditum Netolitzky, 1920. Specie attribuite a Bembidion subg. Ocydromus Clairville, 1806: B. radians Andrewes, 1922; B. charon Andrewes, 1926. Specie attribuite a Bembidion subg. Peryphophila, Netolitzky, 1942: B. endymion Andrewes, 1935. Nuove sinonimie: B. tienmushaniense Kirschenhofer, 1984 = B. perditum Netolitzky, 1920 n.syn. ; B. nanpingense Kirschenhofer, 1984 = B. peleum Jedlitka, 1933 n.syn.; B. niedli Jedlitka, 1965 = B. (Ocydromus) yunnanum Andrewes, 1923 n.syn.; B. kinfushanum Jedlitka, 1958 = B. (Ocydromus) hasurada Andrewes, 1924 n.syn.; B. (Pseudolimnaeum ) jedlickanum Toledano, 1999 = B. (Pseudolimnaeum) arnosti Nakane, 1978 n.syn. Ridescrizioni: B. (Philochtemphanes) Netolitzky, 1942; B. (Philochtemphanes) exquisitum Andrewes, 1923; B. (Philochtemphanes) hansi Jedlitka, 1932; B. (Philochtemphanes) perditum Netolitzky, 1920; B. (Aureoplataphus) Netolitzky, 1942. B. (Aureoplataphus) aureofuscum Bates, 1883. Key words: Coleoptera, Carabidae, Bembidiini, Bembidion, Philochtemphanes, Josefia, Bembidromus, Aureoplataphus, palaearctic region, revision, taxonomy MATERIALS AND METHODS This paper is based on study of about 1000 specimens belonging to most palaearctic subgenera. Sources of material are the collections of the following institutions and specialists: BMNH The Natural History Museum, London CB Coll. Baehr, Miinchen CBL Coll. Bulirsch, Lovosice CBr Coll. Brivio (P.I.M.E.), Monza Systematic notes on the paleartic Bembidion, mainly from China i CE Coll. Echaroux, Dourdan CF Coll. Facchini, Piacenza CFk Coll. Farkac, Praha CM Coll. Morita, Tokyo CMa Coll. Magnani, Cesena CMb Coll. Marggi, Bern CMk Coll. Morvan, Karentoir CN Coll. Neri, Forli CP Coll. Pavesi, Milano CPz Coll. Pütz, Eisenhtittenstadt CS Coll. Schmidt, Milano CSc Coll. Sciaky, Rostock CSk Coll. Schiilke, Berlin (CH Coll. Toledano, Verona CW Coll. Wrase, Berlin NMB Naturhistorisches Museum, Basel NMP Narodni Muzeum, Praha NMW Naturhistorisches Museum, Wien UZMK Universitetes Zoologiske Museum, Kgbenhavn ZSM Zoologische Staatssammlung, Miinchen The measurements, made with a Leica MZ12 stereobinocular microscope at 25 x (body) and 100 x (median lobes of aedeagi), are expressed in the text by these abbreviations: pw/pl pronotum width / pronotum length ratio pw/hw pronotum width / head width ratio el/ew elytral length / elytral width ratio ew/pw elytral width / pronotum width ratio The body length has been measured from front margin of clypeus to the apex of ely- tra, and the antennal length from base of the antennomere | to the apex of 11. The dissections were made using standard techniques; genitalia and small parts were preserved in Euparal, on acetate labels fixed on the same pins as the specimens. The pictures were taken with a Nikon F90x camera on a Leica MZ12 stereobinocu- lar microscope; the drawings were made with a drawing tube on the same microscope, scan- ned and processed with the Color-it!2.3™ program on a Macintosh Performa 6400 com- puter. In the descriptions of the median lobes of aedeagi, the definitions of Belousov & Sokolov (1996) are used for the sclerites; the simplified definitions of Toledano (1998, 1999) are used only for the subg. Philochtemphanes. INTRODUCTION For oriental Bembidiini the supraspecific classification is often problematic: many oriental species are intermediate between subgenera that in the western palaearctic region are easy to separate from each other. The impossibility to include them with certainty in these subgenera sometimes led to the description of new subgenera for few species. However, in most cases the uncertain systematic position of the oriental species suggests that the vali- dity of some subgenera should be re-examined. Andrewes (1921, 1922a, 1922b, 1923, 1924, 1927, 1930, 1932, 1935, 1936), descri- 8 TOLEDANO bed hundreds of species from the Indian region, but never affirmed anything about their subgeneric placement. To simplify the identification he used groups of species without systematic importance, as the author himself admits (Andrewes 1935), based on charac- ters which have only specific value. | To attribute the oriental species of Bembidion to subgenera, and reorganize the syste- matics of the groups, is a very long and difficult work. This task was begun by Müller- Motzfeld (1985, 1988, 1998), and is one of the aims of my research (Toledano & Sciaky, 1998; Toledano, 1999). During the last few years I discovered that some Chinese species I was studying were undescribed. To demonstrate this, I had to compare them with most known taxa occurring in China and in the neighbouring regions (eastern Siberia, India, Central and South-Eastern Asia, Japan, North America). But to understand fully their systematic position is a work requiring a much longer time. As a first step to determining their relationship, I re-exami- ned some supraspecific taxa, and found some characters suitable as supraspecific indica- tors. As a result, the placement of several species into subgenera became clear. After this reexamination of the fauna, some small oriental subgenera became larger. For example, the subg. Philochtemphanes Netolitzky, 1942, here revised, formerly inclu- ded only two species; it now includes five species. The subgenus Aureoplataphus Netolitzky, 1942, here revised as well, was described as monotypic, and now includes two species. For one group of new species very strictly related to each other, I describe a new sub- genus, Josefia n. subg.; Bembidromus n. subg., is described for a single species, very 1so- lated, showing very interesting diagnostic characters. Before dealing with the individual species that will be the subject of this work, some historical notes about the different arrrangements used in the past in the supraspecific syste- matics of the Bembidiini must be given, in particular regarding the limits of the genus Bembidion. | Netolitzky (1942-43), Lindroth (1963, 1976, 1980), Erwin & Kavanaugh (1981), Müller-Motzfeld (1985, 1986a, 1986b, 1998), Maddison (1993) dealt with Bembidion as a very large genus, including almost all the species of Bembidiini. Maddison (1993) correc- tly explained the reasons to keep as independent genera Asaphidion Gozis, 1886, Phrypeus Casey, 1924, Bembidarenas Erwin, 1972 and Zecillenus Lindroth, 1980. In my opinion a few more supraspecific taxa are valid as genera: Ocys Stephens, 1829 (fusion of mentum and submentum, separated by a suture in the rest of the Bembidiini) (Vigna Taglianti, 1993, 1994; Kryzhanovsky et al., 1995); Amerizus Chaudoir, 1868 (habitus similar to Trechini and very long mandibles) inclu- ding species from North America (subg. Amerizus) and from Nepal and southwestern China (subg. Tiruka Andrewes, 1935) (Perrault, 1981, 1985; Deuve, 1998); Orzolina Machado, 1987 from Canary Islands (peculiar chaetotaxy, geographically isolated, apterous, intertidal, containing a single species); Caecidium Uéno, 1971 (blind, depigmented Japanese Bembidiini with habitus extre- mely similar to Trechini and elytra with a transverse furrow in addition to basal peduncle). Perhaps Pseudophilochtus Wollaston, 1877 and Endosomatium Wollaston, 1877 from St. Helena, because of the extreme geographical isolation, could be interpreted as belon- ging to an independent genus. However, as I have examined only the external habitus of a Systematic notes on the paleartic Bembidion, mainly from China 5, few species, which are very peculiar in having extremely rounded elytra, I cannot say yet if a generic separation could be acceptable. One interesting diagnostic character of Pseudophilochtus, the absence of the hind pronotal seta, is shared also by several species from New Zealand (Lindroth 1976) and by the chinese B. (Bembidion) sciakyi Toledano, 1999: in my opinion this is another demon- stration that the genus Bembidion is extremely variable and, therefore, characters that in a relatively restricted geographical area seem to be valid for generic or subgeneric diagno- sis, in the examination of the world fauna can loose their supraspecific importance. Nevertheless most species of the world Bembidiini, more than a thousand, in my opi- nion belong to Bembidion Latreille, 1802, since the species included in the independent genera mentioned above are few. REVISION OF BEMBIDION SUBG. PHILOCHTEMPHANES NETOLITZKY, 1942 SYSTEMATIC NOTES. Before I deal with the species of Philochtemphanes Netolitzky, 1942, a reevaluation of some characters important for the diagnosis of the subgenus is in order. In the opinion of most specialists the Bembidiini are divided into two large groups by the position of the discal elytral pores. According to some authors (e.g. Perrault 1981) this character has a generic value, and it separates, for example, the genus Bembidion from the genus Ocydromus. An interesting new way of observation of this character, suggested to me by my friend Maurizio Pavesi, can demonstrate that this is not a good generic cha- racter. Observing the elytral discal pores from the lower side of the elytron (diaphanized) shows that in most cases in which the discal pores appear to be in the third stria they are actually in the third interval. For example, in Microsinocys Toledano, 1998 the discal pore punctures appear from above to be in the third interval but near stria 3; as seen from beneath they are clearly in the third interval (figs. 1-2). Certainly most species of Ocydromus Clairville, 1806 sensu Kryzhanovskij et al. (1995) have the discal pores near the third ely- tral stria, sometimes exactly in the stria, whereas the species included by Jeannel (1941) in his phyletic lines of Notaphus Stephens, 1829 and Bembidion Latreille, 1802 have the discal elytral pores clearly in the third interval. Therefore, even though this character certainly has a taxonomical importance, and it can be useful for the subgeneric determination, in my opinion it does not have a generic value, being a very variable character also in series of specimens of the same species, collected at the same time and in the same place. For exam- ple, I have observed variation in the placement of the discal pores in B. (Ocydromus) deco- rum Zenker in Panzer, 1801. Therefore I believe that Ocydromus should not be accorded generic rank, and that most species of Bembidiini belong to the very large worldwide genus Bembidion, as mentioned above. Confirmation of this view may come from phylogenetic studies of other characters. For example, the examination of the karyotype (e.g. Serrano & Galian, 1996) seems to confirm the homogeneity of the genus, except for a few taxa, probably really representing independent genera. An example of the latter is Synechostictus Motschulsky, 1864, which also shows many interesting taxonomical characters, suggesting the same conclusion, in the male and female genitalia. In any case, Philochtemphanes cer- tainly belongs to the group of subgenera with the discal pores of elytra in the third inter- 10 TOLEDANO val, and it must be kept as subgenus of Bembidion, as Netolitzky (1942) himself did. For the same reason Andrewes (1935) included B.exquisitum Andrewes, 1923, in his group of B.varium, one of his species groups lacking systematic importance, as mentioned above. Philochtemphanes was described by Netolitzky (1942) for B. exquisitum Andrewes, 1923, and B. hansi Jedlicka, 1932. In his original description of the subgenus, the author notes the absence of the apical stria. As suggested by Andrewes (1935), in B. exquisitum the apical stria, even though clearly reduced, actually is always present. In any case, this character in itself has no supraspecific value: many species clearly not related to each other share this character: for example, Bembidion subg. Omoperyphus, Bembidion (Ocydromus) cordicolle Duval, 1851, many species of Bembidion subg. Nepha, Bembidion (Josefia) n. subg., B. algidum Andrewes, 1935, B. misellum Harold, 1877, etc. The presence of this cha- racter in species of the same subgenus can be due to a common derivation, but the presen- ce in different phylogenetic groups does not necessarily suggest any close systematic rela- tionship. Another character used by Netolitzky (1942) in his original description of the sub- genus is the presence of the "crista clavicularis" (fig. 3). The crista clavicularis is a sharp carina beginning from the end of the basal margin of the elytron, more or less extended in the direction of the articulation of the elytron. This is, in my opinion, an important cha- racter for the supraspecific diagnosis of this subgenus, of course in combination with some others, since it is not peculiar of this group. In fact it is present in some other subgenera with the discal elytral pores in the third interval, Emphanes Motschulsky, 1850, Leja Dejean, 1821, Diplocampa Bedel, 1896, Bembidion Latreille, 1802 s.str., Philochtus Stephens, 1829, subgenera surely related to each other, and in Peryphophila Netolitzky, 1942. But all these subgenera are clearly distinguishable from Philochtemphanes, as described in the following key. KEY TO THE PALAEARCTIC SUBGENERA WITH CRISTA CLAVICULARIS 1 - Elytra markedly widened posteriorly, evidently wider than pronotum, with maximum width behind middle, pronotal median line extended to basal margin; base of pronotum rectilinear, sometimes with sides slightly oblique, never lobate; discal elytral pores in the third interval, adjoining stria PROBEN RR cols ears I ASE La AG EME ed LIKE à; Peryphophila - Elytra not markedly widened posteriorly, more parallel and narrower; pronotum often with base lobate, with a median lobe more or less protruding behind; if base of pronotum not lobate, then with median line not reaching basal margin; discal elytral pores in the third interval, more distant LE EL LIO INR E PI Pea ee AO E OPA 2 2 = Fronial nıtows Verdes, and stone ACONMÉLSONE 2. ie done a mt ter nie dE: Leja Frontal furrows more superficial, not or only slightly convergent ....................... 3 3 - Fourth humeral umbilicate pore more distant from the third than the remainder are from each RA RA asi pre i ae e i e e ei EA eve ER = - Humeral umbilicate pores: closely and regularly distributed . 0.10.00 Wr. sr ee 5 EEE EEE REIT TENERE PR oh Diplocampa one nie rar ne N a er] Emphanes 5 - Base of pronotum emarginate, with a median lobe posteriorly protruding; basal foveae almost IRE a zur DE a act atria el Th th ee Peres Mara Tas | dee iaia dti ES 6 - Base of pronotum rectilinear, sometimes only slightly oblique at sides; basal foveae present, squa- re and gloss; elytra piceous brown to black, with a more or less evident yellow apical spot in most Systematic notes on the paleartic Bembidion, mainly from China a Figs. 1-2. Details of the anterior elytral discal pore of Bembidion (Microsinocys) ginghaicum: 1 - seen from above; 2 - seen from below. The channel of the discal pore is truncated-conical and sligh- tly oblique; the center of its larger base (on the lower end) is more distant from stria 3 than the cen- ter of the smaller base (at the upper end of the channel), therefore in fig. 2 the discal pore is more distant from stria 3 than in fig.1. Fig. 3: “Crista clavicularis” of Bembidion (Philoctemphanes) bran- cuccii n. Sp. SPECIES at re SI EI VIRE ER DEE Sen age Philochtemphanes 6 - Pronotum not cordate, transverse, with basal median lobe more or less strongly protruding behind; at the sides of the lobe marsinurecilineat cool. Lett sxe Genre nun Ad Li Philochtus - Pronotum strongly cordate, narrow, with very short emargination at the sides of the base, with hind angles sharp, less sharp if the ind seta is absent. 22. n Bembidion The male genitalia are rather similar in the different species of the subgenus Philochtemphanes, sharing the presence of two main sclerites in the median lobe of the aedeagus reminiscent of sclerites "a" and "b" of Microsinocys Toledano, 1998 and the simi- lar sclerites "alpha" and "sigma" of Bembidion Latreille, 1802 (Toledano, 1999). In the descriptions of the species they will be called "sclerite a" and "sclerite b". In addition to the species previously included in the subgenus, B. perditum Netolitzky, 1920 shows the combination of characters that in my opinion is peculiar for Philochtemphanes: discal elytral pores in the third interval, crista clavicularis present, four humeral umbilica- te pores closely and regularly distributed, frontal furrows almost parallel and superficial, base of pronotum not (or very slightly) lobate: this species, originally described as belon- ging to the subg. Emphanes, is transferred here into Philochtemphanes. | 12 TOLEDANO Two more species of the subgenus are described here as new. The study of the type material of B.tienmushaniense Kirschenhofer, 1984 (correct name for B. tienmushaniensis Kirschenhofer, 1984) revealed that this species is a junior synonym of B. perditum: | B. tienmushaniense Kirschenhofer, 1984 = B. perditum Netolitzky, 1920 n.syn. In the main Collection of NMP I found a specimen of B. (Philochtemphanes) exqui- situm Andrewes, 1923 labelled by Jedlicka himself as type of an undescribed B. andrewe- si (nec B. (Peryphus) andrewesi Jedlicka, 1932), which probably represents nothing more than a "nomen in collectione". Unfortunately, I could not examine the type material of two species, Bembidion lep- taleum Andrewes, 1922 (from India) and B. goetzi Jedlicka, 1965 (from China), that in my opinion could belong as well to Philochtemphanes. In Lorenz (1998) these species are dealt with as B. (Neja) leptaleum and B. (Peryphanes) goetzi. In the provisional checklist of the chinese Bembidiini provided below I include the latter of these two species, but with doubt. TYPE SPECIES. Bembidion exquisitum Andrewes, 1923 REDESCRIPTION. Small species (3.28 to 4.50 mm) piceous to brown, normally with one api- cal elytral spot at the posterior third of the elytral side, very faint in some specimens of B. perditum (Holotype), and in almost all the type specimens of 5. daliangi n. sp. In the paraty- pe of B. daliangi in coll. Farkac there is a light suggestion of elytral apical spots, even thou- gh very faint and with very shaded edges. Head with almost parallel and superficial frontal furrows, normally developed eyes, rather convex, and antennae short. Pronotum with rather large basal foveae impunctate; base very slightly sinuate at sides, but without a real median lobe protruding posteriorly. Anterior margin almost recti- linear, with anterior angles very slightly protruding from the margin. This character seems to be rather variable within the examined specimens. Lateral channel normally widened posteriorly. Laterobasal carina normally short, but well developed. Median line not deep, not reaching the basal margin. Base punctate in all species except B. brancuccii n. sp: Metasternal process distinctly bordered. Legs rather short. Elytra with rather marked shoulders. Presence of more or less developed crista cla- vicularis at the end of the basal margin. Striae finely punctate, ending before apex except for stria | and 2, normally reaching apex; intervals flat. Stria 7 absent in B. hansi and B. brancuccii n. sp. Stria 8 in the anterior third in the sulcus with four umbilicate humeral pores regularly and closely distributed, then separate from the sulcus, very deep, laterally surrounding a wide carina bounded in the inner side by the apical stria. Two discal pores in the third interval. Apical stria extremely reduced and superficial. Male genitalia. In all species a pair of sclerites very similar to "sclerite a" and "scle- rite b" of Microsinocys Toledano, 1998 and sclerites "alpha" and "sigma" of Bembidion Latreille, 1802 (Toledano, 1999) is present. Three apical setae in both parameres. Female genitalia. Spermatheca (fig.43) without annulus receptaculi and with reser- Systematic notes on the paleartic Bembidion, mainly from China TS voir relatively large, elongate, divided in two cavities, the basal one, larger, "bean shaped", connected at the basal extremity to the duct by a small "funnel-shaped" channel and at the apical extremity with the second cavity, smaller, almost cylindrical. Large linear sper- mathecal gland. DISTRIBUTION (fig. 4). The species of the subgenus inhabit the south-eastern palaearctic region from northern India to southwestern China (Yunnan, Sichuan, Shaanx1) where they seem to prefer habitats at relatively high altitude, from about 2000 to 4000 m. BIOGEOGRAPHICAL CONSIDERATIONS. The fauna of Yunnan shows many elements in com- mon with the south-eastern asian region and northern India, as shown by the distribution of B. exquisitum Andrewes, 1923. But the high endemism rate in the SW China shown by many groups of Bembidiini (Deuve, 1998; Toledano, 1998, 1999; Toledano & Sciaky, 1998) and other Carabidae (Sciaky, 1993, 1994; Sciaky & Facchini, 1997, 1999) and also by B. (Philochtemphanes) hansi JedliCka, 1932, is confirmed and emphasized by the new taxa described here. KEY TO SPECIES 1 - Elytra with more or less evident microsculpture in fine, transverse meshes, at least near apex; apical elytral spots clearly visible, with rather sharp edges; if elytral spots faint, then dark elytra with evident elytral microsculpture; pronotum very slightly sinuate before hind angles (figs. 11, 12:13): Li Pra cle Bn aoe Es a de ni AE A cos Ek, FO tse A 2 - Elytra without microsculpture; if faint elytral microsculpture then isodiametric; apical elytral spots absent or very hardly visible; pronotum more evidently sinuate before the hind angles (figs. 14, 18016, 19 DA Le a RARI LETT Mie es RL LIO RARE Ae en UA + 2 - Microsculpture between eyes very superficial or absent; elytra more deeply punctate; stria 7 pre- sent (Northern India; Nepal; China: Sichuan, Yunnan) ........ B. exquisitum Andrewes, 1923 - Microsculpture between eyes isodiametric, very evident; elytra less deeply punctate; stria 7 absent (China: Yanmar, Sichlian). „1. ere I. eee ee in a 3 3 - Smaller (3.46 to 4.22 mm), base of antennae red (at least antennomere 1), rest of antennae brown; legs red; pronotum without evident microsculpture on disk and with punctured base; elytra rather glossy, reddish brown to dark brown with lighter apex and with striae well impressed (China: nani Siemans; „as a. cds a at ae A e IRE RITO B. hansi Jedlicka, 1932 - Larger (4.40 to 4.50 mm), first antennomere brown, rest of antennae darker; legs dark brown; pronotum with faint microsculpture on disk and with base rugose, not punctate; elytra matt, piceous, with striae more superficial (China: Yunnan) ................. B. brancuccii n. sp. 4 - Pronotum more transverse (pw/pl > 1.40) (fig.17), elytra shorter (China: Sichuan, Yunnan, Shaanxi); apical elytral spot absent, only in one specimen very faint, small ........... B. daliangi n. sp. - Pronotum less transverse (pw/pl < 1.31) (figs. 14, 15, 16), elytra more elongate; large apical ely- tral spot, triangular, sometimes very faint (China: Kiautschau, Zhejiang, Shaanxi)........... ER eee een A B. perditum Netolitzky, 1920 B. (Philochtemphanes) exquisitum Andrewes, 1923 (fig. 5) Bembidion exquisitum Andrewes, 1923 TYPE LOCALITY. India, W. Almora Div., Kumaon, Uttar Pradesh. EXAMINED MATERIAL. Holotype, 2, "W. Almora Div., Kumaon U. P., Oct. ‘19 H. G. C." (BMNH), 14 TOLEDANO 19, Paratype, "W. Almora Diva, Kumaon U.P., May 1917 NGC." (BMNH); 16, Tatsienlu, Prov. Setschwan, China Merid. (BMNH); 56 4,49 2, China, Sichuan, Dayi Dafeishui forest, cca 110 km W of Chengdu, 22.6.1993 (CT, CS); 32 2, China - Sichuan, Mt. Emei, 3100 m, 180 km S of Chengdu, 7 July 1993 (CT); 26 4,68 2, China - W Sichuan, Emei Shan 2800 m, 25-31.V1.92 (CS, CT); 66 6, 4 2, China: Sichuan, Emeishan, 180 km SSW Chengdu, 1700 m, 22.6.1994 (NMW, CT); 14,299, China / Sichuan, 103°20' E, 29°30 N, Mt. Emei, 500-1200 m, 4.-18.V.1989 (NMB); 19, China: Sichuan, Mt. Emei, 600-1050 m, 5.-19.5.1989 (NMB); 19, China, pr. Sichuan, Emei Mt., 2500 m, 4.-20.5.1989 (NMB); 1%, China, pr. Sichuan, Emei mt. 2500 m, 4.-20.5.1989 (NMB); 1d, China / Sichuan, 103.36 el/31nw, Quanxian 700 m, 20.V.1989 (NMB); 15,3% £, China (C Sichuan), Qincheng Shan, NW Chengdu 650-700 m (leafy wood), 30°54' N, 103°33' E, 18.V.1997 (CW, CT); 13, China (C Sichuan), Qincheng Shan, NW Chengdu 1000-1200 m, 30°55' N, 103°35' E, 3.VI.1997 (CW); 19, China: Sichuan (2), Qincheng Shan, rückseite 650-700 m, 30°53'.56 N, 103°33'01 E, 18.5.1997 (CW); 19, China: Sichuan, Qincheng Shan, 65 km NW Chengdu, 103°33' E, 30°53' N,, 18.V./ 3.- 4.VI 1997, 8 km W Taiping, 800-1000 m (CPz); 28 5, 29 £, China - S Sichuan, 27°45' N, 101°13' E, pass 20 km S Muli/Bowa, mixed forest, ca 3500 m (CT, CS); 26 d, China - Sichuan, Baoxing, 100 km N of Yaan, 12-14 July 1995 (CT); 1¢, China - S. Sichuan, Daliang Shan mts., 9.-11. VI. 1998, road Meigu - Leibo vill., pass 15 Km NE of Meigu, 28°26'N, 103.17'E (CT); 684,289, China S. Sichuan, Daliang Shan mts., Zhaojue vill. env. pass Xichang - Meigu vill. 12.-14.VI.1998 (CT); 16, 12, China, S. Sichuan, 7. VII.1998, 27.38N, 102.48E, 10 km SW Butuo, cultural steppe (CT); 12, Szechwan mer., Mts. Kinfushan, 2000 m pr. flum. Sung-Kanho (labelled as type of B. andrewesi Jedlicka in litt. nec B. andrewesi Jedlicka, 1962) (NMP); 19, China: W Sichuan, Aba Tibet Aut. Pref. Weizhou Co., Quionglai Shan, Wolong Tal, 20 km WNW Dujia,gyan, 1100m; 31°05N, 103°26E. Schotter, Blüten, 14. VII.1999 (CW); 32 2, China, Yunnan prov., Gaoligongshan mts., 90 km W of Baoshan, 26-28.5.1995 (CS, CT); 1¢, China: NW Yunnan, Yulonxueshan NP near Baishui, ca 30 km N Lijang, 2800-3200 m, 7-11.7.1994 (NMW); 14,1%, China - Yunnan, 100 km W Kunming, Diaolin nat.res., 22.5-2.6.1993 (NMW, CT); 46 6,72 9, Nepal, Hille, VI 1996 (CT, CS); 16, Yunnan, 2600-3100 m, 25°38' N, 100°09' E, Gangshan mts., 5-6/6/1993 (NMB); 1d, China: N Yunnan, Dali, 1600 - 2000 m, 5.-8.VII.1990 (NMB); 26 ¢, 12, Yunnan, 2800-3000 m, 25°12' N, 100°24' E, Weibaoshan mts., 29-30/6.92 (NMB, CT). DESCRIPTION. Length 3.28 to 4.22 mm. Head and pronotum piceous, elytra brown to piceous with lighter apex, with one rectangular yellow spot at the posterior fourth, extended from about stria 4 to the lateral margin. Sometimes some faint iridescence on pronotum and, more evident, on elytra. Legs reddish with femurs infuscated. Antennomeres 1 to 3 and the basal 3/4 of the antennomere 4 red, rest of antennae slightly darker. Head with parallel frontal furrows not extending to the clypeus. Normally without micro- sculpture between the frontal sulci; in few specimens there is a very superficial micro- sculpture, almost invisible, as isodiametric meshes. Pronotum (fig. 11) transverse (pw/pl = 1.30 to 1.42) with square and smooth basal foveae. Sides slightly sinuated, basal margin with median lobe posteriorly protruding. Base with few punctures. No evident microsculpture on the disk. Legs rather short. Elytra (el/ew = 1.50 to 1.55) (ew/pw = 1.44 to 1.46) rather convex, normally with maximum width slightly behind the middle. Square shoulders. Striae 1 to 2, sometimes 3 reaching the apex, punctate in the anterior 3/4 and sulcate in the posterior fourth. Striae 4 to 6 disappearing at the posterior third. Stria 7 present, visible in the anterior half. Intervals slightly convex and shiny. Apical stria not deep, with pore puncture at middle. Scutellar Systematic notes on the paleartic Bembidion, mainly from China 15 Er su h Nei Mongol : 25 "00f ‘M INDIA — TROP IG OF GANGER-—_ Fig. 4. Map of China showing the distribution of chinese species of the subgenus Philochtemphanes: B. exquisitum (1), B. hansi (2), B. brancuccii (3), B. perditum (4) and B. daliangi (5). striole rather long. Crista clavicularis short. Discal elytral pores clearly in the third inter- val, often the anterior one in a small fovea extended to the interval 3. Elytral microsculpture of very fine transverse meshes, often superficial and present only near apex, sometimes visible on the whole elytra, but in this case hardly visible. Male genitalia (fig. 31). Median lobe of the aedeagus slightly curved ventrally. Sclerite "a" wide, and sclerite "b" divided in three oblique strips, the median one wider than the others and longer than in B. hansi. DISTRIBUTION (fig. 4). N India, Nepal, China: Yunnan, Sichuan. AFFINITIES. Similar to B. hansi and B. brancuccii n. sp. in the general habitus, but more related to B. perditum and B. daliangi n. sp. as suggested by the external structure of the male genitalia; the structure of the inner sac is shared by all Philochtemphanes, with very slight specific differences. B. (Philochtemphanes) hansi Jedlicka, 1932 (fig. 6) Bembidion hansi Jedlicka, 1932 TYPE LOCALITY. China, Yunnan fou. EXAMINED MATERIAL. 1 2, Holotype, "Yunnan fou" (NMP); 3d d, Paratypes, "Yunnan fou" (NMP, NMW); 19, Paratype, "Yunnan fou" (NMW); 26 d, China Prov. Yunnan, Vallis flumin. Soling-ho (NMP); 26 6, Vallis flumin. Soling-ho, Yun. (NMW); 15, 12, China, prov. Yunnan, Vallis flumin. Soling-ho (NMP); 23 4,39 ©, Yunnan, Xishan mts., 24°57' N, 102°38' E, 2300 m, 27/6/1993 (NMB, 16 TOLEDANO CT); 19, Yunnan, Yulong mts., 27°01' N, 100°12' E, 24-26 May 1993 (NMB); 16, China - Yunnan, 28.5 - 9.6. 1994 Dali (CS); 14, China - Yunnan, 28.5.-9.6.1994 Dali (CS); 16, China - S. Sichuan, Daliang Shan mts., road Meigu - Leibo vill., pass 15 Km NE of Meigu, 28°25'N, 103.17'E, 9-11.6.1998 (CT); 28 2, China S. Sichuan, Daliang Shan mts., Zhaojue vill. env. pass Xichang - Meigu vill. 12.- 14.VI.1998 (CT). DESCRIPTION. Length 3.46 to 4.22 mm. Head and pronotum piceous black, elytra reddish brown to dark brown, rather matt, with lighter apex. Elytral spots as in B. exquisitum, but on average extended also on interval 4, and with edges less sharp than in B. exquisitum. Legs red. Antennomere 1 and 2and the base of antennomere 3 red, rest of antennae darker. In some specimens antennomere 2 is also dark. Head with parallel frontal furrows not extending to the clypeus, with coarse micro- sculpture in isodiametric meshes on the whole front. Eyes slightly more convex than in B. exquisitum. | Pronotum (fig. 12) transverse (pw/pl = 1.41 to 1.47) almost identical to that of B. exquisitum, with square and smooth basal foveae. Sides slightly sinuated, basal margin with median lobe slightly less protruding posteriorly than in B. exquisitum. Base with few punc- tures. No evident microsculpture on the disk. Legs rather short. Elytra (el/ew = 1.52 to 1.54) (ew/pw = 1.50 to 1.58) with the same shape as in B. exquisitum but slightly less convex. Striae | to 3 reaching apex, sulcate in the apical fourth, punctate in the rest of the elytra. Striae 4 to 6 disappearing in the posterior fourth. Stria 7 absent. Intervals flat and matt. Crista clavicularis short. Discal elytral pores clearly in the interval 3, in a fovea extended on the whole interval 3, sometimes from interval 2 to 4. Microsculpture in fine transverse meshes, slightly more evident than in B. exquisi- tum. Male genitalia (fig.32). Median lobe of the aedeagus clearly wider from the apical third to the middle than at the base. Sclerite "a" rather narrow, and sclerite "b" divided in three oblique strips, the median one wider than the others. A small tooth at the apex, not sharp. DISTRIBUTION (fig. 4). China: Yunnan, Sichuan. AFFINITIES. Easy to recognize from B. exquisitum by the lighter and rather matt elytra, without elytral stria 7 and the coarse microsculpture in the front, between the eyes. It can be recognized from B. brancuccii n. sp. by the smaller size, the paler appendages, and the less developed elytral microsculpture. Closely related to B. exquisitum and B. brancuccii n. sp., but surely a sister species of the latter. The similarities with B. brancuccii n. sp. are stronger, by the structure of the elytral striae and by the presence of coarse microsculptu- re on the head. Also the examination of male genitalia reveals close systematic relation- ships between these species, sharing a structure of the median lobe of the aedeagus cha- racterized by the maximum width of the lobe apical to the middle. Probably B. brancuccii n. sp. is derived from an isolated population near B. hansi with subsequent increase in the size of the body and male genitalia. Systematic notes on the paleartic Bembidion, mainly from China | 17 Figs. 5-10. Photographs of the habitus of: 5 - B. exquisitum, paratype; 6 - B. hansi, holotype; 7 - B. brancuccii, paratype; 8 - B. perditum, holotype; 9 - B. daliangi, holotype and 10 - B. humerale petri- bulirschi, holotype. B. (Philochtemphanes) brancuccii n. sp. (fig. 7) DIAGNOSIS. A Philochtemphanes larger than other species of the subgenus, very similar to B. hansi but characterized by almost completely dark antennae, pronotum microsculptu- red on the disk and matt elytra with reduced elytral striae. TYPE LOCALITY. N Yunnan, 27°49' N, 99°43' E, cca 3600 m, Zhongdian. 18 TOLEDANO TYPE SERIES: Holotype, ¢, "N Yunnan, 27°49' N, 99°43' E, cca 3600 m, Zhongdian, 19-25.VI.1994" (NMB). Paratypes: 14,29 2, same date and locality as the Holotype (NMB, CT); 19, "Yunnan, 25 Km E of Zhongdian, 3300-4000 m, 12-14 Jul. 1995" (NMB). DERIVATIO NOMINIS. Dedicated to Dr Michel Brancucci of the Naturhistorisches Museum, Basel, who kindly sent me the type material in study. DESCRIPTION. Length 4.40 to 4.50 mm. Piceous black. The lighter paratypes have dark brown elytra with a faint yellow spot at the elytral base. Yellow apical spot very faint, from about interval 4 to the side. Legs piceous brown. Antennomere | and 2 dark brown, rest of antennae piceous. Head with parallel frontal furrows not extending to the clypeus, more superficial than in B. hansi. Coarse isodiametric microsculpture on the whole front. Eyes as in B. hansi. Pronotum (fig. 13) transverse (pw/pl = 1.39) with square and smooth basal foveae, sides slightly sinuated, basal margin with median lobe slightly protruding posteriorly as in B. hansi. Base rugose, without punctures. Rather superficial microsculpture in short, tran- sverse meshes also on the disk, Legs slightly longer than in B. hansi. Elytra (el/ew = 1.50 to 1.53) (ew/pw = 1.55) as in B. hansi, but with more superficial striae, stria 1 to 3 reaching apex, sulcate in the apical fourth, punctate in the rest of the ely- tra. Striae 4 to 6 disappearing in the posterior fourth. Stria 7 absent. Intervals flat and matt. Crista clavicularis more developed than in B. hansi. Discal elytral pores clearly in interval 3, in a fovea extended across the whole interval 3, sometimes from interval 2 to 4, but more superficial than in B. hansi. Microsculpture in fine transverse meshes, shorter and more evident than in the two precedingspecies. Male genitalia (fig. 33). Median lobe of the aedeagus similar than in B. hansi, but clearly larger, and more parallel. DISTRIBUTION (fig. 4). China: Yunnan. AFFINITIES. See under B. hansi. B. (Philochtemphanes) perditum Netolitzky, 1920 (fig. 8) B. (Emphanes) perditum Netolitzky, 1920 Bembidion tienmushaniensis Kirschenhofer, 1984 (correct name B. tienmushaniense) SYSTEMATIC NOTES. This species has been described as belonging to the subg. Emphanes Motschulsky, 1846 only because of the presence of the crista clavicularis, but the humeral elytral pores clearly are not in the characteristic position of the subg. Emphanes. Therefore it is transferred here into Philochtemphanes. The pronotal shape is very variable in the examined specimens of B. perditum; the type specimen (fig. 14) shows less developed sinuosity of the lateral margin, while few specimens from Shaanxi (fig. 16) show the maximum development of this character. As shown in fig. 15, B. tienmushaniense has a degree of sinuosity of the lateral margin inter- mediate between the two forms shown in figs. 14 and 16. The comparison of the male geni- talia reveals that all these forms belong to the same taxon. Systematic notes on the paleartic Bembidion, mainly from China | 19 ,00°° oo, @ È « » @ 6 Figs. 11-23. Pronotal shape of: 11 - B. exquisitum; 12 - B. hansi; 13 - B. brancuccii; 14 - B. perdi- tum, holotype; 15 - B. perditum (holotype of B. tienmushaniense); 16 - B. perditum, extreme of sinuo- sity of the pronotal side; 17 - B. daliangi. 18 - B. belousovi; 19 - B. pieroi; 20 - B. taguense; 21 - B. luisae; 22 - B. shugela; 23 - B. panda. Fig. 24: metasternal process of B. belousovi. 20 TOLEDANO TYPE LOCALITY. China: Kiautschau. EXAMINED MATERIAL. 1%, Holotype, "Kiautschau, China" (NMW); 18, Holotype of B. tienmusha- niense Kirschenhofer, 1984, "China, Tien Mu Shan 30° 23’N, 119° 37’ E, 16. VI. 1937, Eigin Suenson leg." (UZMK); 1 2, paratype of B. tienmushaniense Kirschenhofer, 1984, "China, Tien Mu Shan 30° 23’N, 119° 37’ E, 30. V. 1937, Eigin Suenson leg." (NMW); 26 d, 19, "China, Shaanxi, Qing Ling Shan mts., road Baoji - Taibai vill., pass 35 km S of Baoji, 21-23. VI. 1998" (CT); 19, "China - Shaanxi - Qin Ling Shan 108°47'E, 33°51'N, Mountain W pass at Autoroute km 70, 47 Km S Xian, 2300 - 2500 m sifted, 26-30.08.1995" (CW). DESCRIPTION. Length 3.52 to 3.96 mm. Piceous black, antennomere 1 and 2 and base of 3 and 4 red, rest of antennae darker (in the specimen in CW, rest of antennae very slightly infuscated). Legs reddish, with femurs slightly infuscated. Apical spot triangular, larger than in the other Philochtemphanes, even though with unsharp edges. The type is teneral, thus its colour is completely reddish - brown, only with a faint suggestion of the apical spots. The type specimens of B. tienmushaniense I could observe are the same colour; in the paratype of B. tienmushaniense (NMW) the apical spots are sli- ghtly more evident. | Head with normally convex eyes, frontal furrows parallel, not deep, and front smooth, glossy. Pronotum (fig. 14, 15, 16) (pw/pl = 1.17 to 1.31) more cordiform than in other Philochtemphanes; very variable in shape with the more or less sinuate lateral margins. Basal foveae smaller than in the preceding species. Median line not deep, basal transverse Impression evident, base punctured. Legs rather short. Elytra rather elongate (el/ew = 1.47 to 1.60) (ew/pw = 1.52 to 1.62), almost parallel, with square shoulders. Striae 1 to 7 punctate, clearly impressed, stria 3 to 7 disappearing slightly before apex; crista clavicularis well developed. Scutellar striole rather long. Apical stria short. No microsculpture on head, pronotum and elytra. Male genitalia (fig. 34). Shape of the medium lobe of the aedeagus very similar to B. daliangi n. sp., slightly narrower than in the latter, and with sclerite "b" wider, ovate, and with a strongly sclerotized part at the basal end of sclerite "a". DISTRIBUTION (fig. 4). China: Kiautschau, Zhejiang, Shaanxi. AFFINITIES. Surely related to B. daliangi n. sp. and B. exquisitum as indicated by the structure of the male genitalia. It shares with B. daliangi n. sp. weak development of the apical spots, and absence of elytral microsculpture; this last in my opinion is an important diagnostic character within Philochtemphanes. Also the male genitalia are very similar in both species. B. (Philochtemphanes) daliangi n. sp. (fig. 9) DIAGNOSIS. A Philochtemphanes with short, unspotted (only a very faint spot, difficult to see, in a few specimens) elytra, with maximum width slightly behind the middle. TYPE LOCALITY. China - S. Sichuan, Daliang Shan mts., road Meigu - Leibo vill., pass 15 Km NE of Meigu, 28°25'N, 103.17'E. Systematic notes on the paleartic Bembidion, mainly from China | Dil TYPE SERIES. Holotype, d, "China - S. Sichuan, Daliang Shan mts., road Meigu - Leibo vill., pass 15 Km NE of Meigu, 28°25'N, 103.17'E, 27.VII.1997" (CT); Paratypes: 1d, same date and locality as the holotype (CT); 16, "China - S. Sichuan, 26. VII. 1998, 28°25'N, 103.17'E, road Meigu - Leibo, Daliang Shan mts., vill., pass 15 Km NE Meigu" (CT); 12, "China Songpan, N. Sichuan, 3.6.95 (CF); 16, "China, N Yunnan, Zhongdian env., 3200-3300 m 21.-22.V1.1998 27°50N, 99°36E (CFk); 1g, "China, W.Sichuan (Ganzi Tibet. Aut. Pref. Yajiang Co.), Shalui Shan, river valley 6 km WSW Yajiang 3250 m, 30°01N/100°57E (river bank, bank slope), 4.VII.1999 (CW); 15, "CHINA (Shaanxi) Qin Ling Shan 110.06E, 34.25N, Hua Shan, 118 km E Xian, S. top, 1950 - 2000 m. mix. wood, NO. VIII. 1905 (CW), DERIVATIO NOMINIS. Derived from the type locality. DESCRIPTION. Length 3.48 to 3.64. Brown to piceous-black, with greenish metallic reflec- tions in a few paratypes, legs reddish with infuscated femurs. Antennomeres 1, 2 and base of 3, 4 red, rest of antennae darker. Elytra unspotted; in the paratype in Coll. Farkac a light suggestion of apical spots as in most species of Philochtemphanes, even though very faint and with very shaded edges. Head with front smooth, glossy, with few transverse rugosities in few specimens. Pronotum (fig. 17) cordate, small, more transverse than in the other species (pw/pl = 1.40 to 1.42). Median line strongly impressed, ending before the basal margin, with a small median elongate fovea at its posterior end, more evident than in the few specimens of the other species sharing this character. Laterobasal carina very short. Small and square basal foveae extending onto the basal margin, which is strongly uneven if seen from behind. Legs short. Elytra (el/ew = 1.50 to 1.52) rather short, with maximum width behind the middle and square shoulders. Stria 7 less impressed than the others. Striae disappearing very sli- ghtly before the apex. Scutellar striole rather long. Apical stria very short. Microsculpture absent. In the paratype from Songpan in CF a very superficial iso- diametric microsculpture is present on the elytral apex. Male genitalia (fig. 35). Sclerite "b" sharp, clearly narrower than in B. perditum. DISTRIBUTION (fig. 4). China: Sichuan, Yunnan, Shaanxi. Bembidion subg. Bembidion s.str. Latreille, 1802 Bembidion (Bembidion) humerale petribulirschi n. ssp. (fig. 10) In my revision of subgenus Bembidion (1999) I mentioned a female specimen of B. humerale with completely dark elytra collected in Turkey by my friend Dr. Petr Bulirsch. After examining some more specimens sharing the absence of elytral spots I could confirm that they belong to a new taxon. The reasons that lead me now to describe this taxon as a new subspecies of B. hume- rale Sturm, 1825, are: first, all the specimens of B. humerale collected in Turkey I could examine at present share the absence of elytral spots; second, I have not found in the lite- rature any mention of the typical form of B. humerale from there; third, only one comple- tely black species of Bembidion s.str. is mentioned from Piiliimur, Turkey, B. quadrima- culatum cardiaderum Solsky, 1874 (Korge, 1971), but I suspect that this mention could be 22 TOLEDANO referred to B. humerale petribulirschi n. ssp., because also the typical form of B. quadri- maculatum Linné, 1761, is present in Turkey, and the geographical distribution of B. qua- drimaculatum cardiaderum does not suggest its presence in Turkey (Toledano, 1999). Another reason is that in many cases in the subg. Bembidion geographically isolated popu- lations share a darkening of the integument, with loss of the elytral spots. DIAGNOSIS. A B. humerale from Turkey with completely dark elytra. TYPE LOCALITY. Ulukisla, Anatolia (Turkey) TYPE SERIES. Holotype, 6, "Ulukisla, Anat., 28.VII.47, Exp. N. Mus. CSR" (CFk). Paratypes: 19, "Turkey (Antalya), Saklikent 1600-2100 m, 9/7/1998 Bey Daglari, leg. Bulirsch" (CBL); 1 à , Sivrihisar, Anat., 7.IX.47, Exp. N. Mus. CSR" (CT); 19, "Turkey, prov. Ordu, Koyulhisar, 29.5.1989, leg. Schönmann et Schillhammer" (NMW); 19, "TR K Maras, dint. Goksun, 12/V/1997, leg. Sama” (CN); 19, "TR K Maras, 13 km SGoksun, 12/V/1997, leg. Magnani” (CMa). DERIVATIO NOMINIS. I dedicate this subspecies to my friend Dr. Petr Bulirsch of Lovosice who collected and kindly gave me for study the first specimen of this new taxon I could seg, DESCRIPTION. Length 2.64 to 2.88 mm. Piceous-black, antennomere 1 black, rest of anten- nae dark brown, slightly paler the apex of antennomere 11, femurs and base of tibiae dark, rest of legs yellow, palps dark brown. Elytra completely black. Head rather small, with rather deep and almost parallel frontal furrows, extending to the clypeus, where they are slightly convergent; convex eyes, antennomeres rather short. Pronotum transverse (pw/pl = 1.23 to 1.32) slightly wider than head (pw/hw = 1.03 to 1.09). Elytra rather elongate (el/ew = 1.60-1.63) and parallel, with well developed shoul- ders. Striae pointed, rather superficial, visible in the basal 5/6. Stria 1 pointed in the basal 2/3 and sulcate in the apical third. Male and female genitalia identical to those of the nominotypical form. DISTRIBUTION. Turkey. This is the southernmost occurrence of B. humerale. AFFINITIES. This species lacks the apical tooth of the median lobe of the aedeagus. As jud- ged by this character, B. humerale has a relatively independent position within the Bembidion sensu stricto: the lack of the tooth is shared only with B. crassicorne Putzeys, 1872, a spe- cies otherwise not strictly related to B. humerale. B. humerale petribulirschi is easy to reco- gnize from some other taxa of the subgenus sharing the absence of elytral spots by the black antennomere 1, since B. quadrimaculatum ssp. cardiaderum Solsky, 1874 (Turkmenistan) and B. quadrimaculatum ssp. caporiaccoi Netolitzky, 1934 (N India) have always the base of the antennae more or less red, never black, while B. quadripustulatum Serville, 1821, always with at least antennomere 1 black and clearly larger in all its subspecies, present in the same region of B. humerale ssp. petribulirschi, never shows completely black elytra. The remaining palaearctic Bembidion species without elytral spots, B. sciakyi ssp. luguen- se Toledano, 1999 from China, sharing also the black first antennomere , is clearly distin- guishable by the absence of the seta at the hind pronotal angle, and by the apical structure of the median lobe of the aedeagus. My key to the palaearctic species and subspecies of subgenus Bembidion (Toledano, 1999) has to be slightly corrected by adding the new subspecies described here at couplet 9: Systematic notes on the paleartic Bembidion, mainly from China | 23 9 Ebtraicompletet dar Turkey st Bela) Toni ot humerale petribulirschi n. ssp. -., Blytracwith-at least one pale spot, rot. sunto Ir 2 wa dgl: 10 Bembidion subg. Josefia n. subg. DIAGNOSIS. A subgenus of Bembidion characterized by small size (3.26 to 4.40 mm), anten- nae moniliform, head with very superficial frontal furrows; pronotum depressed and cor- _ date, distinctly sinuate at sides before the hind angles, with extremely superficial basal foveae; elytra ovate with striae more or less lightly punctured, discal pores in the third inter- val, near stria 3, and apical elytral pore isolated. TYPE SPECIES. Bembidion belousovi herewith described. DERIVATIO NOMINIS. The subgenus is dedicated to my friend Dr. Josef Jelinek of the Narodni Muzeum of Praha; the gender of the name is feminine. DESCRIPTION. Small species (3.26 to 4.40 mm). Piceous-black with legs reddish to brown. Head relatively large with normally convex eyes; antennae moniliform as in Microsinocys Toledano, 1998, antennomeres slightly more elongate only in B. belousovi. Frontal furrows superficial, not extending to the clypeus, rarely extending posteriad behind the middle of the eye, sometimes extremely reduced, almost absent. Pronotum cordate, depressed, with sides more or less sinuate before the hind angles. Basal foveae extremely superficial, flat, with lateral carina more or less evident, never com- pletely developed. Base of pronotum rectilinear, sometimes slightly sinuate at sides. Lateral anterior pore puncture about at the anterior third, posterior pore puncture at the corner of the hind angle. Metasternal process (fig.24) with a large border, with a transverse furrow at the midd- le of the mesocoxae. Legs rather short in all species. Elytra strongly oval, with rounded humera. Elytral striae normally superficial, punc- tate. In one species (B. pieroi n. sp.) striae more deeply punctate, in B. luisae n. sp. and B. shugelae n. sp. striae punctate - sulcate. Only stria 1 reaching the apex, joining there the lateral sulcus. Stria 8 in the anterior third in the lateral sulcus, with four humeral pores clo- sely and regularly distributed, then faint, but visible, sulcate, separate from the sulcus, ending in line with the apical pore, isolated by the absence of the apical stria. Scutellar striole short. Basal elytral margin ending with a small tooth between the beginning of striae 5 and 6 (at the beginning of stria 6 in B. luisae and B. shugela). Anterior discal elytral pore in the third interval, almost on stria 3, the posterior one slightly more distant from stria 3. Male genitalia with a well-sclerotized ribbon brush, not large, in the median lobe of the aedeagus; in all the species except for B. luisae n. sp. a basal chitin platelet is present. Median lobe of the aedeagus ventrally curved in all species. Parameres with 3 setae. Female genitalia (fig.44). Spermatheca with small reservoir almost triangular and a well sclerotized and a long annulus receptaculi which is "trumpet shaped". DISTRIBUTION (fig. 46). Southwestern China (NW Sichuan, Qinghai, C Tibet). AFFINITIES. The subgenus Josefia groups species very closely related to each other. It belongs to the group of subgenera of Bembidion with discal elytral pores in the third interval. Within 24 TOLEDANO this group Josefia seems to be rather isolated. The common character of having the apical elytral pore isolated, as discussed above, probably due to inheritance from a common ance- stor, does not mean any close relationship with other subgenera sharing the absence of the apical stria. The group that seems to be most closely related is the chinese subg. Microsinocys Toledano, 1998, as indicated by the structure of the head and the antennomeres, the colour and the small size of its species. The differences that reveal distinction between Josefia and Microsinocys at a subge- neric level are in the structure of the pronotum, sinuated at sides (not sinuated at sides and often with a tooth at the hind angles in Microsinocys), in the elytral structure, with elytral striae slightly impressed, but evident, and apical pore isolated (elytra almost completely flat with striae evanescent and with carinated apical stria in Microsinocys). In addition there are differences in male genitalia, with the internal sac of the median lobe of the aedea- gus in almost all species of Josefia one of the two typical large sclerites of Microsinocys is missing; the curved shape of the median lobe of the aedeagus in Josefia is clearly reco- gnizable from the more rectilinear shape of Microsinocys. KEY TO SPECIES 1 - Pronotum smaller (ew/pw = 1.34 to 1.46), strongly cordate, with base clearly narrower than the anterior margin (figs. 18, 19, 20); laterobasal carina of pronotum rudimentary but present; spe- Ciesayithout evigent elytral mierosceulpture (er belousovi)t cirie ale ee ann 2 - Pronotum larger (ew/pw = 1.31 to 1.37), less cordate, with base not narrower than the anterior margin (figs. 21, 22); laterobasal carina of pronotum absent; elytral microsculpture present in one SU AL PEN EN ARTE L'ART CUT TAR EL RE EEE RER ci 2 = tlytral giniae more superficiali pronotiim more any assi Le. ne, aa ia 3 - Elytral striae with deeper punctures, pronotum more depressed (fig. 19) ......... pieroi n. sp. Oo 1 Larger size (3.84 to 3.96 mm); pronotum slightly less transverse (pw/pl = 1.39); elytral stria 7 normally absent or very superficial; median lobe of the aedeagus as in fig. 36; Rongbaca Da le LE) NEE, EIN a RL RE a, belousovi n. sp. - Smaller size (3.52 to 3.62 mm); pronotum slightly more transverse (pw/pl = 1.42 to 1.43)); ely- tral stria 7 evident; median lobe of the aedeagus as in fig. 38; Sichuan ........ taguense n. Sp. 4 - Pronotum more sinuate at sides, before hind angles (fig.21); elytral microsculpture evident, in fiausverseneshes(IS Sichuan. E Dinehal), = su. ile oto luisae n. Sp. - Pronotum less sinuate at sides (fig. 22), before hind angles; elytral microsculpture completely ATIC USA ILI, a Ra AC) EN atte FRA Sr ar e shugela n. sp. gr. belousovi B. (Josefia) belousovi n. sp. (fig. 25) DIAGNOSIS. The largest species of the subgenus Josefia, with transverse pronotum. TYPE LOCALITY. China NW Sichuan, 31.46N 99.32E, pass 20 km NW Rongbaca, Picea forest 4091m. TYPE SERIES: Holotype, d, "Ch. NW Sichuan, 31.46N 99.32E, pass 20 km NW Rongbaca, Picea fore- st 4091 m, 20.7.1995" (CT). Paratypes: 17d d, 109 2, same date and locality as the holotype (CT). DERIVATIO NOMINIS. I dedicate this species to my friend Dr Igor Belousov of St. Petersburg, Russia, well known specialist of Carabidae and in particular of Bembidion. Systematic notes on the paleartic Bembidion, mainly from China 25 Figs 25-30. Photographs of the habitus of: 25 - B. belousovi, holotype; 26 - B. pieroi, paratype; 27 - B. taguense, holotype; 28 - B. luisae, holotype; 29 - B. shugela, holotype; 30 - B. schuelkei, holotype. DESCRIPTION. Length 3.84 to 3.96 mm. Piceous black, glossy, with faint metallic greenish reflections. Sides of elytra slightly lighter than the rest of elytra. Legs red-brown, anten- nomere |, 2 and base of 3, 4 red, rest of antennae piceous. 26 TOLEDANO Head relatively large, glossy and convex front with superficial and parallel frontal furrows, not extending to the clypeus; eyes normally convex. Antennae short with anten- nomeres slightly more elongate than in the other species. Pronotum (fig. 18) transverse (pw/pl = 1.39) (ew/pw = 1.40) with well rounded mar- gins, not strongly sinuated behind, before the hind angles, sharp, square. Base narrower than the anterior margin. Basal margin slightly sinuate, with lateral angles slightly more advanced than the median part of the margin. Basal foveae small, triangular, rather deep. Rudimentary laterobasal carina, almost invisible in some specimens, base evidently punc- tate with basal transverse impression well impressed. Median line evident, not deep. Elytra (el/ew = 1.45 to 1.46) oval, strongly convex. Basal margin reaching a point between the beginning of stria 5 and 6. Elytral striae finely punctate, stria 7 slightly less impressed than the others, almost absent in most specimens, striae 2 to 7 ending slightly before the apex. Intervals flat. No microsculpture on the whole body, except for the super- ficial isodiametric meshes of the neck. Male genitalia (fig. 36). Median lobe of the aedeagus larger than in the other species, very wide. Well sclerotized ribbon brush, not large, basal and ventral chitin platelet rather evident and developed. Basal part of the median lobe strongly curved to the left side. DISTRIBUTION (fig. 46). Known only from the type locality in China, Sichuan. AFFINITIES. Within Josefia this species forms a species group together with B. taguense n. sp.and B. pieroi n. sp. as indicated by the structure of the inner sac of the median lobe of the aedeagus and the pronotal shape, with base narrower than anterior margin. B. belouso- vi differs from these species only by the larger size, the more convex pronotum and the male genitalia. From B. pieroi n. sp. it is also recognizable by the less impressed elytral striae. B. (Josefia) pieroi n. sp. (fig. 26) DrAGNosis. The species of Josefia with transverse pronotum with coarser punctuation of the elytra. TYPE LOCALITY. China, W Sichuan, 3000 m Litang. TYPE SERIES: Holotype, ¢, "China, W Sichuan, 3000 m Litang, 3-7-94" (CS). Paratypes: 19, same date and locality as the holotype (CS); 2d d, "China, SW Sichuan, r. Xiangcheng - Derong, p. 10 km NW Xiangcheng, 14-7-94" (CF, CT); 19, "China Litang, 10 Km South m. 4000, Sichuan 9. VIT.1992" (CS). DERIVATIO NOMINIS. Dedicated to Dr. Piero Toledano, my dear brother and my colleague in the dental practice. DESCRIPTION. Length 3.26 to 3.50 mm. Colour as in B. belousovi. Head, compared with B. belousovi, with frontal furrows slightly sharper, even thou- gh not deep, and front less convex. Antennae moniliform, shorter than in B. belousovi. Pronotum (fig. 19) transverse (pw/pl = 1.38 to 1.40) (ew/pw = 1.34 to 1.45) with base evidently narrower than the anterior margin. Sides of pronotum anteriorly rounded, strongly restricted behind, oblique and very slightly sinuate before the hind angles, small and obtuse. In the hind half, lateral channel very deep. Anterior angle slightly protruding anteriorly from Systematic notes on the paleartic Bembidion, mainly from China 24 0 43 4 4 45 mm Figs. 31-42. Left view of the median lobe of the aedeagus of: 31 - B. exquisitum; 32 - B. hansi; 33 - B. brancuccii; 34 - B. perditum; 35 - B. daliangi; 36 - B. belousovi; 37 - B. pieroi; 38 - B. taguense; 39 - B. luisae; 40 - B. schuelkei; 41 - B. panda; 42 - B. aureofuscum. Fig. 43-45: reservoir of the sper- matheca of: 43 - B. exquisitum; 44 - B. belousovi; 45 - B. jaechi. 28 TOLEDANO the anterior margin. Median line very slightly impressed, not reaching the basal margin; basal transverse impression deep. Base very slightly punctate, with large but superficial punctures. Laterobasal carina evident, sharp, well developed in almost all the specimens. Elytra (el/ew = 1.44 to 1.47) oval. Basal margin reaching a point between the begin- ning of stria 5 and 6. Elytral striae 1 to 7 coarsely punctate, disappearing slightly before apex. Only stria 1, sulcate in the apical fourth, is deep to apex where it reaches stria 9. Intervals more convex than in B. belousovi. Apical pore isolated. No microsculpture on the whole body, except for the superficial isodiametric meshes of the neck. Male genitalia (fig. 37). Median lobe of the aedeagus narrower and with the apex more angulate ventrally than in B. belousovi, with the same sclerites. DISTRIBUTION (fig. 46). China: Sichuan. AFFINITIES. Belongs with B. belousovi and B. taguense to a species group within the subg. Josefia, characterized by the structure of the pronotum, more transverse than in the group of B. luisae. Within this group it is more closely related to the other relatively small spe- cies, B. taguense, as shown by the male genitalia, and it can be recognized from B. taguen- se by the pronotal shape and the coarser elytral punctuation. B. (Josefia) taguense n. sp. (fig. 27) TYPE LOCALITY. Cina - W Sichuan, Tagu 3700 m. TYPE SERIES. Holotype, d, "Cina - W Sichuan, Tagu 3700 m, 27.VII.92" (CS); paratypes: 24 4,29 2 same date and locality as the holotype (CS, CT); 12, "China, SW Sichuan, r. Xiangcheng - Derong, p. 10 km NW Xiangcheng, 14-7-94" (CS); 29 2, "China W-Sichuan (Ganzi Tibet Aut. Pref. Litang Co.), Shalui Shan, 25 km NW Litang, 4200 m, 30°08N/100°04E, 1.-3.VII.1999 (CW, CT); 16,19, "CHINA: W-Sichuan Ganzi Tibet. Aut. Pref., Litang Co. Shalui Shan, FluBufer 25 km NW Litang, 4200m, 30°08N, 100°04E, Schotter, Steine, 1. VII. 1999" (CW); 18,32 2 "CHINA: Prov. Sichuan Ganzi Tibetan Auton. Pref., Batang Co., Shalui Shan, Abies-Forest, 55km NE Batang, 30.07.68N, 100.03.77E, 4200m, 3.VIL.1999" (CPz, CT). I labelled as "Bembidion subg. Josefia cfr. taguense" one female specimen from "China, SW Sichuan, road Litang - Sumdo, pass 50 km SSW of Litang, 5.7.94" (CS), because it differs from the others by the more superficial striae and the more depressed pronotum. It seems similar to B. taguen- se, but may be an hybrid with B. pieroi; in any case I prefer to not include it in the type series. DERIVATIO NOMINIS. Derived from the type locality. DESCRIPTION. Length 3.52 to 3.62 mm. Colours as in B. belousovi. Head with front almost completely flat, glossy, with frontal furrows parallel, extremely superficial, with posterior end more advanced than in B. belousovi and B. pieroi. Antennae moniliform, slightly more elongate than in B. pieroi. Pronotum (fig. 20) transverse (pw/pl = 1.42 to 1.43) (ew/pw = 1.43 to 1.46), very similar to that of B. belousovi, though less convex, with base narrower than the anterior margin and hind angles as in B. belousovi. Laterobasal carina sharp, well developed. Transverse basal impression deep, faintly punctate. Basal foveae triangular, rather deep. Elytra (el/ew = 1.43 to 1.45) oval, slightly more depressed than in B. belousovi. Basal margin reaching a point between the beginning of stria 5 and 6. Striae 2 to 7 very slightly Systematic notes on the paleartic Bembidion, mainly from China 29 Nei Mongol; _| ALU e - maura = ang ibet) : 2500 7002, TROPIC OF CANCER Fig. 46. Map of southwestern China showing the distribution of B. belousovi (1), B. pieroi (2), B. taguense (3), B. luisae (4), B. shugela (5), B. schuelkei (6) and B. panda (7). more deeply impressed than in B. belousovi, ending slightly before apex. Stria 7 present, even though more superficial than the others and ending at the apical 2/3, more developed in few specimens, where it reaches apex with few punctures. Apical pore isolated. No microsculpture on the whole body, except for the superficial isodiametric meshes of the neck. Male genitalia (fig. 38). Median lobe of the aedeagus similar to B. pieroi, but sligh- tly wider and with the apex less angulate ventrally. Same sclerites in the internal sac as in B. belousovi and B. pieroi. DISTRIBUTION (fig. 46). China: Sichuan. AFFINITIES. See under B. belousovi and B. pieroi. er. luisae B. (Josefia) luisae n. sp. (fig. 28) DIAGNOSIS. A Josefia with pronotum clearly larger and less transverse than in the other spe- cles. TYPE LOCALITY. China, NW Sichuan, 32.30 N 98.25 E, pass 20 km S Quagca. TYPE SERIES. Holotype, d, "Ch. NW Sichuan, 32.30 N 98.25 E, 17-18/7/1995, pass 20 km S Quagca, alpine meadow, + 4100 M" (CT); paratypes: 1d, "China, E Qinghai prov., Qingshuihe, 4200 m, 1.- 30 TOLEDANO 5.7.1992" (CS); 38 &, "China - W Sichuan, r. Litang - Batang, p. 70 km WNW Litang, 4675 m, 22.7.94, alp. region" (CE, CS). | DERIVATIO NOMINIS. Dedicated to my beloved mother, Luisa Roncari Toledano. DESCRIPTION. Length 3.44 to 4.40 mm. Piceous black with very faint metallic reflections. Legs dark brown with base of tibiae darker, piceous; basal half of antennomere | and api- cal half of antennomere 2, 3 and 11 dark red, rest of antennae darker, dark brown to piceous. Head flat and glossy with frontal furrows parallel, rather deep, and front with some wide transverse rugosities between the frontal sulci in some specimens. Eyes more convex than in the other Josefia, antennae moniliform. Pronotum (fig. 21) large, rather depressed, less transverse than in the preceding spe- cies (pw/pl = 1.27 to 1.32), with base larger than the anterior margin, with sides rounded in the anterior 4/5, then rectilinear, ending in square hind angles. Base almost rectilinear. Anterior angles not evidently protruding anteriorly. Median line rather deep. Basal tran- sverse impression rather superficial, base impunctate. Laterobasal carina almost absent. Basal foveae very superficial, flat. Elytra (el/ew = 1.50 to 1.68) slightly wider than pronotum (ew/pw = 1.31 to 1.32), oval, but with shoulders more developed than in the precedingspecies. Basal margin rea- ching the beginning of stria 6. Elytral striae rather well impressed, punctate - sulcate, lon- ger than in the precedingspecies, ending very slightly before the apex. Stria 7 present, sli- ghtly less impressed than the others. Apical pore isolated. Microsculpture isodiametric, superficial on the frons, superficial in rather wide, sli- ghtly transverse meshes on the pronotal sides, superficial, in rather large, transverse meshes on elytra. Male genitalia (fig. 39). Median lobe of aedeagus strongly curved ventrally, with nar- row apex. Two main sclerites, on the right side one similar to sclerite "a" of Microsinocys Toledano, 1998, on the left side another one, with two oval small bags toward basal ope- ning and a long narrow tube ending toward apex with an hook. DISTRIBUTION (fig. 46). China: Sichuan, Qinghai. AFFINITIES. Apparently forming together with B. shugela n. sp. a species group within Josefia as indicated by structure of the pronotum. Unfortunately the impossibility of comparing male genitalia does not allow confirmation of this hypothesis. B. (Josefia) shugela n. sp. (fig. 29) DIAGNOSIS. A Josefia very similar to B. luisae, from which it differs by the shape of the pronotum, less sinuate at sides, before the hind angles, and absence of elytral microsculp- ture. TYPE LOCALITY. China - C. Tibet, Shugela Pass, 5100 m, 40 km E of Yambajing. TYPE SERIES Holotype © , "China - C. Tibet, Shugela Pass, 5100 m, 40 km E of Yambajing, 31.5.1997" (CT). DERIVATIO NOMINIS. Derived from the type locality, used in apposition. DESCRIPTION. Length 3.48 mm. Head and pronotum piceous-black with faint greenish metal- lic reflections, elytra and pronotal basal foveae brown (type specimen not completely matu- Systematic notes on the paleartic Bembidion, mainly from China 31 re). Antennomeres 1, 2 base of 3, 4 and apex of 11 red, rest of antennae darker. Legs reddish. Head with superficial frontal furrows extending to the clypeal lateral pores, front glos- sy, slightly rugose between eyes. Eyes convex as in B. luisae. Pronotum (fig.22) (pw/pl = 1.27) (ew/pw = 1.37) similar to that of B. luisae, but with side less sinuate, and more oblique in the hind fifth, where it reaches the basal margin for- ming a slightly obtuse hind angle. Basal margin very slightly curved, base very weakly punctate, basal transverse impression superficial. Median line deeper than in B. luisae. Anterior pronotal angles as in B. luisae. Elytra (el/ew = 1.51) as in B. luisae, only with stria 8 very superficial, faintly punc- tate, and a very superficial apical stria only in the right elytron. No microsculpture on head, pronotum and elytra. Male genitalia. Unknown. DISTRIBUTION (fig.46). Known only from the type locality. SYSTEMATIC NOTES. This is a single female specimen, as is the other mentioned above as "Bembidion subg. Josefia cfr. taguense", but in this case the strong difference in the shape of the pronotum and the geographic distance suggests that this is a species distinct from B. luisae. AFFINITIES. See under B. luisae. Bembidion subg. Microsinocys Toledano, 1998 A new species of this subgenus is described here. The homogeneity of this subgenus is shown also by this species, which shares almost completely the external shape with B. schillhameri Toledano, 1998, and the median lobe of the aedeagus with B. facchinii Toledano, 1998. The key to the species of Microsinocys in Toledano (1998) must be changed at cou- plet 9, and there must be added a new couplet 11, as follows: 9 - Smaller (2.6 - 2.8 mm) with laterally projecting tooth at hind angles of pronotum; basal margin of pronotum with median lobe evidently protruding posteriorly ....................... ll - Larger (2.9 mm), without laterally projecting tooth at hind angles of pronotum; median lobe of aedeagus large (0.68 mm) with large sclerites and evident ventral gibbosity ................ LE ea Ea ut dalle 6 VITRO RS RA Ie na jani Toledano, 1998 11 -Median lobe of the aedeagus without evident ventral gibbosity and without dorsal emargination; Yunnan, Yulenexuesha ro sus sure di ee eee eee schillhammeri Toledano, 1998 - Median lobe of the aedeagus with evident ventral gibbosity (fig. 41) and dorsal emargination; Sichuan. Maxie Sieam, see e eo schuelkei n. sp. Bembidion (Microsinocys) schuelkei n. sp. (fig. 30) DIAGNOSIS. A Microsinocys with elytra on the average slightly more convex than in B. schil- Ihammeri Toledano, 1998, but almost impossible to distinguish from this species by the external characters. The only character allowing to separate them is the median lobe of the aedeagus, very similar to that of B. facchinii Toledano, 1998. 32 | TOLEDANO TYPE LOCALITY. China: W-Sichuan, Ganzi Tibet. Aut. Pref., Kangding Co., Daxue Shan, 10 km S Kangding, 29.59 N, 101.55 E, 3150 m. TYPE SERIES. Holotype, & , "China: W-Sichuan, Ganzi Tibet. Aut. Pref., Kangding Co., Daxue Shan, 10 km S Kangding, 29.59 N, 101.55 E, 3150 m, Flußtal, Gesiebe, Felsnieschen, 26.VI.1999" (CW). Paratypes: 156 4,39 ©, same date and locality as the holotype (CSk, CW, CT); 146 8,6% 9 "China: W-Sichuan, Ganzi Tibet Auton. Pref. Daxue Shan, 10km S Kangding, 3150m, 29.59.N, 101.55.E, 26.VI.1999 (CPz, CT); 22 2, "CHINA: W Sichuan, 15 km W Kangding, Rie.138, 3250m, 29°57N 102°54E, 19.VII.98" (CMb). DERIVATIO NOMINIS. Dedicated to my friend Michael Schiilke from Berlin, in whose col- lection I found some paratypes of this new species. DESCRIPTION. Length 2.80 to 2.84 mm. Piceous-black, shiny; legs, antennomeres 1, 2 and base of the antennomere 3 brown. Rest of antennae darker. Head as in B. schillhammeri. Pronotum (pw/pl = 1.45 to 1.50) similar to that of B. schillhammeri, with lateral, rec- tilinear, part of the posterior margin slightly shorter than in B. schillhammeri. Elytra (el/ew = 1.30 to 1.33) slightly more convex than in B. schillhammeri on ave- rage. In a few specimens the elytra seem identical. The elytral striae are slightly more impressed than in B. schillhammeri. Male genitalia (fig. 40). Median lobe of the aedeagus showing the typical structure of Microsinocys, with the sclerites "a" and "b"; extremely similar to the aedeagus of B. facchinii. DISTRIBUTION (fig. 46). Known only from the type locality in China, Sichuan. AFFINITIES. Belonging to the schillhammeri group of the subg. Microsinocys and surely derived from a common ancestor with B. schillhammeri, since in my opinion the structu- re of the pronotum in this group is the more important infrasubgeneric character. Bembidion subg. Bembidromus n. subg. For a single very strange species, B. panda n. sp., I describe here a new subgenus, as it does not belong in any known subgenus. DIAGNOSIS. Subgenus characterized by small size (2.76 to 3.04 mm), very large and con- vex eyes, supernumerary setae on the last abdominal sternites, metallic integument and almost complete elytral striae. TYPE SPECIES. Bembidion panda herewith described. DERIVATIO NOMINIS. The peculiar combination of characters shared with subgenera not clo- sely related to each other led me to use for this new supraspecific taxon a name composed of parts of the names of two very important supraspecific taxa of the Bembidiini; the gen- der of the name is masculine. DESCRIPTION. Length 2.76 to 3.04 mm. Integument with faint metallic reflections. Head large, with convex eyes. Front with frontal furrows deep and parallel. | Pronotum with sides very slightly sinuated before hind angles. Basal margin narrower than the anterior one. Front angles evidently protruding anteriorly. Base with a median lobe protruding behind. Sides of the base with an emargination rather deep before the hind angle, Systematic notes on the paleartic Bembidion, mainly from China 33 that, therefore, is acute. Base rugose, impunctate. Metasternal process slightly bordered. Last abdominal sternites with short supernu- merary setae. Legs short. Elytra short (el/ew = 1.36 to 1.38), depressed, with deeply impressed striae, almost complete. Basal margin reaching the beginning of stria 5 where it is curved behind to join the stria with an obtuse angle. Apical stria long and deep, connected to stria 5. Scutellar striole short. Discal elytral pores in the third interval, very near stria 3. Male genitalia (fig. 41). Base of median lobe of aedeagus strongly curved to left side, basal opening rather large, on the right side. Internal sac feebly sclerotized, with a single group of small sclerites. Left paramere with four apical setae, right one with three. Female genitalia. Unknown. DISTRIBUTION (fig. 46). China, Sichuan. AFFINITIES. The peculiar structure of the pronotum, with hind angles posteriorly protruding, does not suggest any close relationship with other subgenera; I am otherwise aware of it, in a more developed form, only in B. (Thaumatoperyphus) ajmonis Netolitzky, 1934, an indian species evidently not related to B. panda n. sp. I believe that in the case of B. panda it could be interpreted as a specific character. The extremely small size and the position of the discal pore punctures, in the third interval near stria 3, the anterior one more evidently in the inter- val, seem to place this subgenus relatively near to subgenera related to Bembidion s.str., as Microsinocys, even though I have previously expressed doubts about the validity of the posi- tion of the discal pore punctures as an indicator of relationships. The structure of the male genitalia does not show any similarity with Microsinocys. The new subgenus shares the ely- tral structure, having almost complete striae, with subgenera clearly not related to Bembidion and Microsinocys, including the group of Plataphus s.1. and Bembidionetolitzkya, but a close relationship with these subgenera is excluded by the structure of the male genitalia. The head with large and very convex eyes is shared by B. (Plataphodes) farkaci Toledano & Sciaky, 1998, occurring in the same region, sharing also the almost complete striae and the metal- lic integument, but all the species of Plataphodes have the basal margin reaching stria 4 while in Bembidromus the basal margin reaches stria 5. The most important diagnostic character of this subgenus, the presence of supernu- merary setae on the last abdominal sternites, is shared with the subgenera Trichoplataphus Netolitzky, 1914, Blepharoplataphus Netolitzky, 1920, and Aureoplataphus Netolitzky, 1942 (see below). The presence of this character in Bembidromus could be matter of con- vergence, not necessarily meaning any close systematic relationship with these subgene- ra, even though the new subgenus shows some similarities in the structure of the internal sac with the species of Blepharoplataphus. In any case the systematic independence of this subgenus is paradoxically emphasized by the sharing of characters with many supraspecific taxa not closely related to each other. Bembidion (Bembidromus) panda n. sp. (fig. 47) DIAGNOSIS. A Bembidion species very small (2.76 to 3.04 mm) showing depressed prono- tum with emarginated basal margin at the hind angles (fig. 24) and almost complete, sul- 34 TOLEDANO cate, elytral striae, and elytral discal pores adjoining stria 3, each one in a fovea extended to interval 3, and therefore belonging to Bembidromus. TYPE LOCALITY. China, W. Sichuan (Aba Tibet. Aut. Pref. Weizhou Co.), Quionglai Shan, Wolong valley, 69 km WSW Dujiangyan, 3900 m, 30°54N, 102°59E. TYPE SERIES. Holotype, à , "China, W. Sichuan (Aba Tibet. Aut. Pref. Weizhou Co.), Quionglai Shan, Wolong valley, 69 km WSW Dujiangyan, 3900 m, 30°54N, 102°59E (alpine meadow), 15. VII.1999" (CW); Paratype, d, same date and locality as the holotype (CT). DERIVATIO NOMINIS. Derived from the common name of Ailuropoda melanoleuca David, 1869, another very interesting (not entomological!) species inhabiting the same region as the species described here. DESCRIPTION. Length 2.76 to 3.04 mm. Black with faint greenish metallic reflections. Legs dark brown with darker femurs. Antennomere | reddish brown, rest of antennae piceous brown. Head large, with convex eyes, narrower than the pronotum. Front smooth with fron- tal furrows deep and parallel, not extending to the clypeus. Pronotum (fig. 23) transverse (pw/pl = 1.48 to 1.51) with sides very slightly sinua- ted before hind angles. Basal margin narrower than the anterior one. Front angles eviden- tly protruding anteriorly. Base with a median lobe protruding behind. The sides of the base with an emargination rather deep before the hind angle, that, therefore, is acute. Base rugo- se, impunctate. Legs short. Elytra short (el/ew = 1.36 to 1.38), depressed, with striae 1 to 5 deep, sulcate, rea- ching apex. Stria 6 and 7, slightly less deeply impressed, ending a little before apex. Basal margin reaching the beginning of stria 5 where it is curved behind to join the stria with an obtuse angle. Apical stria long and deep, connected to stria 5. Scutellar striole short. Discal elytral pores in the third interval near stria 3, the anterior one more evidently in the inter- val, in a fovea extended in the paratype over interval 3 and in the inner half of interval 4, smaller in the holotype. Microsculpture in isodiametric meshes on the neck, in large transverse meshes on the elytra where is faint, and on the pronotum, where is very superficial, rather difficult to see, almost absent. Male genitalia (fig. 41). Median lobe of aedeagus small, linear, with dorsal margin gently curved and rounded apex. Base strongly curved to the left side, basal opening rather large, on the right side. Internal sac poorly sclerotized, with a single group of small scleri- tes, rather compact if seen from the left side, clearly formed by two sclerites, the larger one on the right side, if seen from the ventral side; a very small basal chitin platelet is also visi- ble. DISTRIBUTION (fig.46). Known only from the type locality in China, Sichuan. Bembidion subgg .Trichoplataphus Netolitzky, 1914, Blepharoplataphus Netolitzky, 1920 and Aureoplataphus Netolitzky, 1942 Among the unidentified Chinese material of the Naturhistorisches Museum, Wien, I discovered a taxon of the group of Plataphus sensu lato with supernumerary setae on the Systematic notes on the paleartic Bembidion, mainly from China 35 51 \ è & Figs. 47-52. Photographs of the habitus of 47 - B. panda, holotype (magnification a); 48 - B. aureo- fuscum; 49 - B. jaechi, holotype (magnification b); 50 - B. baehri, paratype; 51 - B. echarouxi, holoty- pe; 52 - B. maddisoni, holotype (magnification c). abdominal sternites, appearantly new, slightly related to the known Bembidion species belonging to the subgenus Trichoplataphus Netolitzky, 1914, which also share this cha- racter. In order to identify the systematic position of this strange species with light red legs, a character unusual for the group, I examined many type specimens of Bembidiini from 36 TOLEDANO China in the Netolitzky's Collection, preserved in the same Museum. During an interesting discussion about the fauna of this area, Dr. Manfred Jach of the NMW suggested that I exa- mine the possibility of systematic relationships with Japanese taxa. The study of a paraty- pe and one specimen of B. (Aureoplataphus) aureofuscum Bates, 1883, in the NMW revea- led that it was related to the new species I was studying. SYSTEMATIC DISCUSSION. Some of the characters used for the subgeneric diagnosis by Netolitzky (1942) are here interpreted as specific, and the diagnosis is here slightly chan- ged to include the common characters of both species. But, before I discuss the subg. Aureoplataphus, I re-examination of the subgenera of Bembidion with supernumerary setae on the abdominal sternites is in order. In the past, some of these subgenera have been synonymized with Trichoplataphus Netolitzky, 1914. In Miiller-Motzfeld (1985) B. kara Andrewes, 1921, the type species of subg. Triporus Andrewes, 1921, is attributed to Trichoplataphus. The synonymy of both subgenera can certainly be accepted because the species of Triporus (B. kara, Andrewes, 1921, and B.tambra Andrewes, 1923) differ from the species of Trichoplataphus only by the presence of three discal setae on the elytra. The number of discal elytral pores in Bembidiini is not necessarily a supraspecific indicator. For example, in Ocys Stephens, 1829 some species having only one discal elytral pore, such as Ocys (Ocys) harpaloides Serville, 1821, and other species having two pores, such as Ocys (Ocys) quinquestriatus Gyllenhal, 1910, certainly belong to the same subgenus. Also in the material I examined recently I could see a specimen of B. (Ocydromus) hasurada Andrewes, 1924, from China, Sichuan, with two discal pores in the left elytron, and three in the right one. The species of Blepharoplataphus Netolitzky, 1920, (e.g. B. hasti Sahlberg, 1827, B. virens, Gyllenhal, 1827, B. hiogoense Bates, 1873) have been separated from Trichoplataphus by the taxonomists because of the length of the abdominal supernumerary setae, a charac- ter not important, in my opinion, and not useful for the taxonomist, since in specimens lacking setae it cannot be detected. For this reason Blepharoplataphus has been listed as synonym of Trichoplataphus by some authors (Jedlicka 1965; Kryzhanovskij et al., 1995). But the valid diagnostic character to isolate the subg. Blepharoplataphus is in my opinion the general body shape, very similar to the habitus of Plataphus s.str. as B. prasinum Duftschmid, 1812, B. gebleri Gebler, 1833, etc., with pronotum rather small and large ely- tra, together with the presence of the supernumerary setae, and the male genitalia, which is very similar in all of the species. Therefore the species of this group are easy to reco- gnize, and they are clearly more related to each other than to species of Trichoplataphus s.str. Therefore I prefer to remove here the subg. Blepharoplataphus from synonymy with Trichoplataphus: Blepharoplataphus Netolitzky, 1920 bon. subg. nec syn. of Trichoplataphus Netolitzky, 1914. Since within the whole group of Plataphus sensu lato red appendages is extremely rare, being a peculiarity of the two species dealt with here below, their independence at a supraspecific level in my opinion is not unlikely, and the validity of the subgenus Aureoplataphus can be confirmed here, even though the colour of legs is seldom a supra- specific indicator among Bembidiini. Systematic notes on the paleartic Bembidion, mainly from China 37 I believe that Trichoplataphus, Blepharoplataphus and Aureoplataphus can derive from a common ancestor near Plataphus s.str., evolved in three lines which independently acquired supernumerary setae. This in my opinion is suggested by the homogeneity of the species within each one of the three subgenera dealt with here. Even though I believe that a very close systematic relationship between the three sub- genera dealt with here and Bembidromus is unlikely, the following key, to be complete, includes also this subgenus. KEY TO THE PALAEARCTIC SUBGENERA WITH SUPERNUMERARY SETAE ON THE ABDOMINAL STERNITES 1- Size smaller; if size of 4.16 to 4.68 mm, then legs red; if legs dark, then very small size (2.76 to 3.04 mm), very convex eyes and pronotum with hind angles protruding posteriorly ..... 2 size larger (more than AH mm) lecedarker: <i shown. ss NR SCONTO eee ei 3 2- Legs red, size larger (4.16 to 4.68 mm), eyes normally convex, pronotum with basal margin gently emarginate at sides and hind angles not protruding behind ...................... riad ale oan es. le tte Miss subg. Aureoplataphus Netolitzky, 1942 - Legs dark, size smaller (about 3.00 mm), eyes markedly convex, pronotum with hind angles protidinepostenondie ss ae ee Sn UE subg. Bembidromus nov. 3- Habitus similar to Plataphus, with metallic, rather matt integument, pronotum rather small, elytra rather wide, with maximum width behind middle; all species with two discal elytral pore a a Ai ela nn ikon ra SSSR ES subg. Blepharoplataphus Netolitzky, 1920 - Species often with piceous - black color, shiny, often iridescent, with pronotum on the avera- ge larger and elytra more parallel, more elongate; some species with three discal elytral pores Be tale re AER SEES DE EEE FOA NARA subg. Trichoplataphus Netolitzky, 1914 REVISION OF BEMBIDION SUBG. AUREOPLATAPHUS NETOLITZKY, 1942 Bembidion (Aureoplataphus) Netolitzky, 1942 TYPE SPECIES. Bembidion aureofuscum Bates, 1883 REDESCRIPTION. Length 4.16 to 4.68 mm. Head, pronotum and elytra metallic with bronze - greenish reflections. Appendages completely light red. Head slightly narrower than pronotum (pw/hw = 1.10 to 1.24) with deep, parallel, furrows, extended to the clypeus, anterior supraorbital pore with a very large umbilicate pore. Eyes prominent and antennomeres rather elongate. Pronotum transverse, rather depressed, with basal foveae rather superficial, square, and well developed laterobasal carina. Anterior transverse impression and median impres- sion superficial, this last not reaching the basal margin. Basal transverse impression more deep. Base impunctate. Legs rather short. Elytra with parallel sides, square shoulders, basal margin reaching stria 5. Umbilicate pore at the common beginning of striae 1, 2 and scutellar. Elytral striae punctate-sulcate. Scutellar stria well developed. Apical stria as a prolongation of stria 5, slightly more deep than the rest of stria. Two discal pores, the anterior one at one third, the posterior one at the two-thirds of elytra. Metasternal process unbordered. 38 TOLEDANO Male genitalia (fig. 42). Known only in the type species. Median lobe of aedeagus with ventral margin rectilinear and apex ventrally bent. Dorsal margin emarginated, with the typical shape of Plataphus s.l. Base strongly bent to the left side. Left paramere appa- rently with four apical setae, right one with three. Female genitalia (fig. 45). Spermatheca simple, with a sclerotized reservoir appa- rently formed by a single cavity. The duct is almost invisible in the holotype of B. jaechi. A poorly sclerotized, small, "bell shaped" annulus receptaculi is present. DISTRIBUTION. Eastern palaearctic region, Japan and China (Hunan). AFFINITIES. Belonging to the group of subgenera near Plataphus with supernumerary setae on the last abdominal sternites. Within this group, it is the subgenus more strictly related to Plataphus as indicated by the structure of the median lobe of the aedeagus. KEY TO SPECIES: 1 - Elytral striae more impressed, stria 7 deeply impressed, pronotum sinuate before hind angles, basal margin of pronotum with emarginate sides (Japan) ........ B. aureofuscum Bates, 1883 - Elytral striae lightly impressed, stria 6 more or less impressed, stria 7 absent, pronotum less sinua- te before hind angles, basal margin of pronotum rectilinear (China) .......... B. jaechi n. sp. B. (Aureoplataphus) aureofuscum Bates, 1883 (fig. 48) TYPE LOCALITY. Japan, Nagasaki, Kumamoto, Miyanoshita (Jedlicka, 1965). Type in the Natural History Museum, London. EXAMINED MATERIAL. 1 6 , Paratype, "Nagasaki, 22.V. - 3.VI.81" "Japan, G. Lewis, 1910-320" "B. aureofuscum, cotype, Bates" (NMW); 14 , "Nippon moyen, Gallois 1912, Mus. Paris dedit" (NMW). REDESCRIPTION. Length 4.16 mm. Elytra bronze, metallic, with faint greenish reflections. Antennomere 1 red, slightly infuscated at apex, rest of antennae dark brown. Legs red. Palps brown, with penultimate article of maxillary palps piceous. Pronotum transverse (pw/pl = 1.32 to 1.36), depressed, with basal margin emargina- te laterally. Sides of pronotum slightly but evidently sinuate before hind angles. Basal foveae rather deep, square, with lateral carina, median line rather superficial, not reaching the basal margin, anterior transverse impression rather shallow, basal transverse impression deeper. Elytra (el/ew = 1.61 to 1.64 mm) with parallel sides, square shoulders, basal margin reaching stria 5, and terminating here with a very small tooth in the paratype, lacking in the other examined specimen. Discal pore punctures adjoining stria 3, foveate. All elytral striae well impressed, complete. Microsculpture rather evident, isodiametric, on head, pronotum and elytra. Male genitalia (fig. 42). Median lobe of the aedeagus with ventral margin rectilinear and apex bent ventrally. Dorsal margin emarginated, with the typical shape of Plataphus s.1. Base strongly bent to the left side. Apex "axe shaped" with rounded extremity. One small sclerite near basal opening. Left paramere apparently with four apical setae, right one with three. DISTRIBUTION. Japan. Systematic notes on the paleartic Bembidion, mainly from China 99 B. (Aureoplataphus) jaechi sp.n. (fig. 49) TYPE LOCALITY. China, NW - Hunan Wulingyuan N Davong, Suoxiyu. TYPE MATERIAL. Holotype, 2, "CHINA, NW - Hunan Wulingyuan N Davong, Suoxiyu, 31.10.1993, 450 m" (NMW); Paratype, £, same data of the Holotype (CT). DERIVATIO NOMINIS. I dedicate this species to my friend Dr. Manfred Jach of the Naturhistorisches Museum, Wien, who suggested to me the way to solve my doubts regar- ding the systematic position of this species. DESCRIPTION. Length 4.60 to 4.68 mm. Dark metallic green with rather matt elytra. Appendages red, except for the penultimate article of maxillary palpi, the apical two-thirds of the anten- nomeres 2 to 4, and the antennomeres 5 to 11, which are dark brown. Pronotum transverse (pw/pl = 1.41 to 1.49), depressed, with rectilinear basal margin, sides slightly sinuate before the hind angles. Basal foveae square, laterally delimited by a carina. Lateral margin a little wider than in B. aureofuscum. Anterior and posterior tran- sversal impression rather evident. Median line rather deep. Elytra (el/ew = 1.47 to 1.55) more flat than in B. aureofuscum, with less impressed striae. Stria 7 absent, stria 6 almost absent in the paratype. The rest of striae reaching the apex, the outer slightly less impressed apically. Discal pores adjoining stria 3, smaller than in B. aureofuscum, not foveate. Microsculpture less evident than in the precedingspecies, isodiametric, in large meshes, on the head, very superficial, in sharp, transverse meshes on the pronotum, isodiametric on the basal impressions of the pronotum, almost invisible on elytra. Male genitalia. Unknown. Female genitalia (fig.45). Spermatheca simple, with a sclerotized reservoir apparen- tly formed by a single cavity. The duct is almost invisible in the holotype. A poorly scle- rotized, small, "bell shaped" annulus receptaculi is present. DISTRIBUTION (fig. 92). China: Hunan. Bembidion subg. Ocydromus Clairville, 1806 An interesting way to deal with the very difficult Ocydromus complex has been pro- posed by Kryzhanovskij et al. (1995): several subgenera of Bembidion have been down- graded to species groups and included in a very large subg. Ocydromus together with many other species groups. These subgenera are: Peryphus Stephens, 1829, Asioperyphus Vysoky, 1986, Synechoperyphus Netolitzky, 1943, Ocydromus Clairville, 1806, Ocyturanes Miiller- Motzfeld, 1986 and Peryphanes Jeannel, 1941. The systematic position of these taxa has been fixed in the past in several different ways. Andrewes (1935) included most Indian species of the Bembidiini in the genus Bembidion, except for the few species belonging to Asaphidion Gozis, 1886, and Tiruka Andrewes, 1935. The Indian species of Bembidion were divided in species groups without systematic importance, as mentioned above, to make the identification of the species easier. Jeannel (1941) included in the genus Peryphus, independent from the genus Bembidion, the French species of the subgg. Peryphus and Peryphanes, together with the species of Nepha Motschulsky, 1864, Daniela Netolitzky, 1910 (= Bembidionetolitzkya Strand, 1929), 40 TOLEDANO Peryphiolus Jeannel, 1941, Omoperyphus Netolitzky, 1931, Testediolum Ganglbauer, 1892 and Euperyphus Jeannel, 1941. Netolitzky (1942-43) kept the palearctic species of this group within Peryphus, as a subgenus of Bembidion, with Peryphus divided into many species groups; Daniela, Nepha, Omoperyphus, Synechoperyphus, Euperyphus and Testediolum are dealt with as indepen- dent subgenera. Antoine (1955), was the first to use the name Ocydromus as the generic name for the groups belonging to Peryphus according to Jeannel (1941), being the oldest valid name for the group, and included within the genus Ocydromus the Moroccan species belonging to the subgenera Peryphus, Peryphanes, Omoperyphus, Nepha and Synechostictus; Daniela is dealt with as synonym of Peryphus. Lindroth (1963) for the Northern American fauna arranged the Bembidion species as belonging to species groups of this genus, except for Phrypeus Casey, 1924, treated as independent genus. Jedlicka (1965) included the eastern palaearctic species within the subg. Peryphus, and divided it into four species groups: gr.ovale, gr. thermarum, gr. poppii and gr. Peryphus (sic!). Perrault (1981) ranged Ocydromus as a genus independent from Bembidion, attribu- ting to the latter the species belonging, according to Jeannel (1941) to the "séries phyléti- ques de Bembidion et Notaphus". According to Perrault (1981) the valid genera of the Bembidiini are: Asaphidion, Amerizus, Synechostictus, Cillenus, Ocys, Lymneops, Odontium, Phyla, Metallina, Bembidarenas, Ocydromus and Bembidion. Müller-Motzfeld (1984, 1985, 1986a, 1986b, 1988) used the name Ocydromus as a subgenus of Bembidion, for the species of the groups decorum, saxatile, atlanticum, and modestum. In the checklist of italian Carabidae, Vigna Taglianti (1993) followed the arrange- ment of Perrault (1981). | Morita (1994b) arranged Plataphus as subgenus of the genus Ocydromus. Miiller-Motzfeld (1998) provided a list of the subgenera of Bembidion near Ocydromus, and described two new subgenera, Politophanes and Terminophanes: the subgenera listed in this work were interpreted as species groups of the subgenus Ocydromus. They are: Synechoperyphus (syn. of Ocydromus s.str.), Peryphanes, Ocyturanes, Peryphiolus, Peryphus, Asioperyphus and Euperyphus. Most species groups mentioned in Müller-Motzfeld (1998) and Kryzhanovskij et al. (1995) have subgeneric names: this is incorrect, since the names of genus rank must not be used for infrasubgeneric groups. Nevertheless, except for this nomenclatorial mistake this way to deal with the Ocydromus complex is in my opinion the best one. I discussed above the character of the position of the discal elytral pores, used to separate both taxa at the gene- ric level by Perrault (1981), a character too inconstant to be used for the generic diagnosis. Moreover, a generic division between Bembidion and Ocydromus adds some pro- blems because it does not allow us to include with certainty several taxa (e.g. Plataphus, Plataphodes, Trichoplataphus, Blepharoplataphus, Aureoplataphus, Parataphus, Hirmoplataphus, Bembidionetolitzkya, Nepha,) within Bembidion or Ocydromus, and an isolation of these taxa in independent genera, according to Jeannel (1941) and Vigna Taglianti Systematic notes on the paleartic Bembidion, mainly from China 41 (1993) splits at too high a level a very homogeneous group as Bembidion sensu lato. To divide into several subgenera, more easy to study, the species groups included by Kryzhanovskij et al. (1995) in the subg. Ocydromus has been one of the aims of the supra- specific study of Bembidiini in the past, as shown above. In my opinion this division is at present also impossible. In fact most groups of species of Ocydromus share several dia- gnostic characters in different combinations, and this often does not allow the separation into monophyletic groups which could be treated as independent subgenera. Therefore I agree with Kryzhanovskij et al. (1995): all these taxa must be attributed to the extremely large and worldwide subg. Ocydromus, divided in species groups. Nevertheless, in this work I prefer to refrain from a formal synonymization of the taxa downgraded to species groups waiting for more evidence. Several of these species groups will be discussed below. Bembidion (Ocydromus) gr. lenae DIAGNOSIS. An eastern palaearctic and nearctic group of species of the subg. Ocydromus characterized by dark integument with unspotted and convex elytra, large and convex pro- notum with basal border wider than the anterior one, head more or less punctate behind the frontal sulci, antennomere 1 red and long main sclerite with a coil in the median lobe of the aedeagus. SYSTEMATIC NOTES. The difficulty in finding of a clear boundary between this and other groups in the complex of Ocydromus sensu lato did not lead any author to isolate the spe- cies of the /enae group into an independent subgenus, even though this need is felt by other specialists (Seiji Morita, personal communication). Miiller-Motzfeld (1998) described a new subgenus, Politophanes, for B. polites Andrewes, 1935 (fig. 56). This species shows a very strange feature in the median lobe of aedeagus: a sclerotized tooth near the apex, on the dorsal side (fig. 59). But the rest of the inner sac of the aedeagus, characterized by a long main sclerite with a coil (present for example also in many species of the lunatum group and, without coil, in many species of Ocydromus s.str. and of the dalmatinum group), and the external shape, are extremely similar to that of the species of the /enae group. I believe that the curious structure mentioned above may have only specific value, and that the contorted main sclerite has more systematic importance. In my opinion B. polites, within its own species group, may be included into the subg. Ocydromus. One japanese species of the lenae group, B. gotoense Habu, 1977, is strictly related to the chinese species B. peleum Jedlicka, 1933, and to the new B. schoenmanni n. sp. Thanks to the kind help of my friend Seiji Morita of Tokyo, I could examine a specimen of B. gotoense, unfortunately not completely mature (fig. 54). The study of this specimen and the comparison of the male genitalia of B. peleum with the drawing of the median lobe of the aedeagus (Habu 1977) confirm my opinion about the strict relationships with B. peleum. In order to check the validity of the new 5. schoenmanni n. sp. I compared it with the type specimens of 5. peleum and B. nanpingense. From this comparison it became evident that these taxa are synonyms: 42 TOLEDANO . B. nanpingense Kirschenhofer, 1984 (correct name for B. nanpingensis Kirschenhofer, 1984) = B. (Ocydromus) peleum Jedlicka, 1933 n.syn. EXAMINED MATERIAL. B. peleum: Holotype, £, "Tatsienlu, Prov. Setschuan, China Merid." (NMP); 16, paratype, "Tatsienlu, Yüling Süd, Szechuan China" (NMP); 16; holotype of B. nanpingense, "China, Nanping 26° 50’N, 118° 05’ E, 17. V. 1935, Eigin Suenson leg." (UZMK); 14 , paratype of B. nanpingense, same date and locality as the holotype of B. nanpingense, (NMW); 85 3, 62 &, China - Shaanxi, Lüeang - 33°07'N, 106°05'E, 22.5-29.5.97 (CF, CKu, CT); 36 6, 32 2 China (C Sichuan), Qincheng Shan, NW Chengdu, 600 m (river bank), 33.55 N / 103.30 E, 3./4.VI.1997 (CT, CW); 46 8,42 9, China Guangxi, 10 km Liuzhou, 160-200 m, (18) 11.11.1993 (NMW, CT); 36 d, 32 2 China c., Xiexian, 111.6' - 34,8', 20km NE Yuncheng, 25.V.1996 (CS, CT). B. gotoense: 13, Tomie, Fukuejima, Goto is., Nagasaki Pref. Japan, 20.1V.1976 (CM). B. polites: 53 3, Phul Choki, 2000 m, Nepal, 20.IX.1983 (NMB, CS, CT). Bembidion (Ocydromus) schoenmanni n. sp. (fig. 55) DIAGNOSIS. A chinese Ocydromus of the lenae group extremely similar to B. peleum with elytra more depressed. Examination of the genitalia is required to distinguish these spe- cies; on average, B. schoenmanni is of slightly larger size. TYPE LOCALITY. China: NW Yunnan, ca. 100 km NW Liyang, Hangduanshan, Jiduan - Weixi, Jiduan Ludie, 2200 m. TYPE SERIES. Holotype, d, "China: NW Yunnan, ca. 100 km NW Lijang, Hangduanshan, Jiduan - Weixi, Jiduan Ludie, 2200 m (10), 1.7.1994" (NMW); paratypes: 46 d, 42 2, same date and loca- lity as the holotype (NMW, CT). DERIVATIO NOMINIS. Dedicated to my friend Dr. Heinrich Schönmann, curator of the Naturhistorisches Museum, Wien, who kindly gave me the type series to study, and who always helps me in my studies by giving in loan the abundant material preserved in his institution. DESCRIPTION. Length 4.52 to 6.00 mm. Black, metallic, with greenish reflections. Antennomere 1 and apical third of antennomere 2 red, apex of antennomere 11 red-brown, rest of anten- nae piceous. Legs dark brown. Head with eyes convex, parallel and corrugated frontal furrows, few very fine punc- tures between eyes, mainly in the posterior part of the front. Antennomeres rather elongate. Pronotum (pw/pl = 1.31 to 1.32) rather depressed with basal margin slightly oblique at sides. Median line well impressed. Base abundantly punctured between the basal foveae, Square, corrugated and not deep. Lateral carina of the hind angle short, but evident. Elytra (el/ew = 1.53 to 1.55) (ew/pw = 1.45 to 1.59) more depressed than in B. peleum Jedlicka, 1933, with square shoulders. Maximum width slightly behind the middle. Striae 1 to 7 well impressed, more superficial in the posterior fourth. Scutellar stria long. Apical stria not deep, joined with stria 5. Intervals flat, except for the first, slightly raised. Discal elytral pores on stria 3, the anterior one at the anterior third, the posterior one at the hind third. Male genitalia (fig.57). Similar to that of B. peleum but with the apex clearly nar- rower, without the dorsal apical gibbosity. Inner sac slightly protruding from the base. Left 43 , mainly from China WLU € on i Systematic notes on the paleartic Bemb de. wir rt NL Pret ren ag I; 54 - B. gotoense; 55 - B. schoenmann 53 - B. peleum; Figs. 53-56. Photographs of the habitus of 56 - B. polites. 44 TOLEDANO paramere with four setae, right one with three. DISTRIBUTION (fig. 92). Known only from the type locality in China, Yunnan. AFFINITIES. Within the lenae group certainly more closely related to B. peleum Jedlicka, 1933 as indicated by the structure of the body and male genitalia. Bembidion (Ocydromus) gr. baehri DIAGNOSIS. Group of alpine Ocydromus species from the southeastern palaearctic region, piceous-brown to black, metallic, characterized by head with eyes flat and rather small, antennae rather short, frontal furrows parallel and superficial, few superficial punctures behind frontal furrows, pronotum rather depressed with basal foveae small, triangular, hind pronotal angles square, without lateral carina, elytra more or less oval, with maximum width behind the middle, humeral angle of elytra rounded, elytral striae more superficial at the apex of elytra, basal margin reaching a point between the beginning of stria 5 and 6, api- cal stria strongly reduced, or absent with isolated apical pore, isodiametric elytral micro- sculpture, more or less developed, legs short, inner sac slightly protruding from the basal opening of the median lobe of the aedeagus and spermatheca simple, with "bean shaped" reservoir, long duct slightly sclerotized and without a strongly sclerotized annulus recep- taculi. SYSTEMATIC NOTES. In the subgenus Ocydromus sensu Kryzhanovskij et al. (1995) several palaearctic species living on high mountains seem to be related to each other. But to find clear evidence of systematic relationships is sometimes very difficult. The shared charac- ters, such as the depressed elytra, reduced wings (and, therefore, rounded humera), piceous to black color, suggest an evolutionary convergence, since the peculiar environment, cold and windy, with dark soil, can favour matt and dark species that do not fly. Also the flat- ness of the eyes, rather small, is a character often present in non-flying species. Sometimes, some of these similar species inhabit mountains rather close to each other, and the possi- bility of systematic relationships is high, as, for example, in the species of the western palaearctic tauricum group Müller, 1918 (Maurizio Pavesi, personal communication). Two species give a striking example of convergence. One is B. alticola Fiori, 1903, an endemic species of Abruzzo, Italy. In Vigna Taglianti (1993) this species is included in Ocydromus subg Peryphanes Jeannel, 1941, but probably this species must be included together with B. reiseri Apfelbeck, 1902, in a species group within the subgenus Ocydromus (Maurizio Pavesi, personal communication). The other species, B. echarouxi n. sp. (fig. 61) from Southwestern China, is almost identical to B. alticola, differing only by the shorter anten- nomeres and the median lobe of the aedeagus, with the inner sac slightly protruding from the base. In my opinion a close systematic relationship between these species is unlikely, and the similarity is only matter of convergence. In all species of the baehri group the inner sac of the aedeagus protrudes slightly from the basal opening: this is a character occurring in some species of Ocydromus in the dal- matinum group, in the lenae group, and in the terminale group. Systematic notes on the paleartic Bembidion, mainly from China 45 Figs. 57-62. Left view of the median lobe of the aedeagus of: 57 - B. schoenmanni; 58 - B. peleum and 59 - B. polites; 60 - B. baehri; 61 - B. echarouxi and 62 - B. maddisoni. Fig. 63: reservoir of sper- matheca of B. echarouxi. KEY TO SPECIES 1 - Size smaller (3.56 to 3.74 mm); elytra more parallel; pronotum less sinuate before hind angles; median lobe of aedeagus asin fig.60 Li. 24, obama ste be Lis cala ehr. 50, - Size larger (4.24 to 4.48 mm); elytra with maximum width clearly behind the middle; pronotum more sinuate before hind angles; median lobe of aedeagus as in fig. 61,62 ...............2 2 - Elytra wider (el/ew = 1.45 to 1.46), strongly oval; antennae shorter; median lobe of aedeagus as in fig. 61; Sichuan, ‘Qinghai, Tibet NU fas em ARI ae 1 a ee OEE. ED, 46 TOLEDANO - Elytra more elongate (el/ew = 1.50 to 1.53), antennae more elongate; median lobe of aedeagus sino 62 a bel ee n pores PR treads cit DOO maddisoni n. sp. B. (Ocydromus) baehri n. sp. (fig. 50) DIAGNOSIS. An Ocydromus of the baehri group smaller than the others, with elytral sides almost parallel and relatively narrow median lobe of the aedeagus. TYPE LOCALITY. Sumdschiling, W. Tibet. TYPE SERIES. Holotype, d, "Sumdschiling, Ebene W. Tibet, Süssw. Tümp, VII/06, Zugm." (ZSM); paratypes: 16 6 d, 19 £ P, same date and locality as the holotype (ZSM, CBM, CT);4 66,799, "China - N Tibet, Tangula Shan Mts., Tangula Pass, 5100-5300 m, 8.-11.7.1997"(CT). DERIVATIO NOMINIS. I dedicate this species to my friend Dr. Martin Baehr, curator of the Zoologische Staatssammlung, Miinchen, who kindly helps me by trying to teach me how to deal with very large groups, such as the Bembidiini. DESCRIPTION. Length 3.56 to 3.74 mm. Specimens from the type locality reddish-brown to dark brown with bronze metallic reflections, with reddish brown legs, antennae reddish brown with base of antennomere 3 and 4 red. Specimens from Tangula Shan darker, piceous- black with greenish metallic reflections, with legs and antennae brown to dark brown, base of antennomere 3 and 4 red. Head with eyes flat and smaller than in the other species of the group, frontal furrows parallel and superficial, few superficial punctures behind frontal furrows. Antennomeres relatively elongate. Legs rather short. Pronotum (pw/pl = 1.28 to 1.39) rather depressed with basal foveae small, rather deep, triangular, hind pronotal angles square, without lateral carina. Median impression very superficial. Basal transverse impression deeper, with base finely punctate. Elytra rather parallel and elongate (el/ew = 1.59 to 1.61) with rounded shoulders; maximum elytral width very slightly behind middle. Elytral striae 1 to 7 visible, punctu- red, stria 7 more superficial, all striae more superficial at apex of elytra, basal margin rea- ching a point between the beginning of stria 5 and 6; apical stria absent, apical pore isola- ted. Intervals rather convex. Microsculpture in isodiametric meshes, coarse on the neck, very superficial at the sides of pronotum and rather evident on elytra. As in all species of the baehri group, except for B. echarouxi, even though the elytral microsculpture is rather evident, the elytra are shiny. Male genitalia (fig. 60). Median lobe of the aedeagus rather narrow with long main sclerite, ribbon brush strongly sclerotized, and large basal chitin platelet flat, translucent, with strongly cheratinized edges in the ventral part. Basal part of the main sclerite and base of the other mentioned sclerites slightly protruding from the basal opening. Left paramere with four setae, right one with three. DISTRIBUTION (fig. 92). China: Tibet. B. (Ocydromus) echarouxi n. sp. (fig. 51) DIAGNOSIS. A relatively large Ocydromus of the baehri group with strongly ovate elytra and wide median lobe of the aedeagus. Systematic notes on the paleartic Bembidion, mainly from China 47 TYPE LOCALITY. E Tibet "Tuntala Shan" pass, road Markam - Zogang, 20 km NE Zogang, 5000 m, 29°43' N, 98°02' E TYPE SERIES. Holotype, d, "E Tibet, "Tuntala Shan" pass, road Markam - Zogang, 20 km NE Zogang, 5000 m, 29°43'N, 98°02'E, alpine meadow, 30.VI-2.VII.1997" (CT); paratypes: 15,32 2, same date and locality as the holotype (CT); 22 2, "Ch-NW Sichuan, 31.56 N 98.53 E, 19.7.1995, Chola Shan, alp. meadow, Chola pass 4600 m" (CT); 14,1%, paratype, "China, E. Qinghai, Nanshan, 7/95" (CE); 533,59 2, "China, Qinghai prov., 300 km SW Xining, Ngola pass, H=4500-4800 m, 10.VII.1999 (CE, CT); "China - NW Sichuan, 4000 - 4500 m, road Garzé - Barong, pass 25 km 2 Garzé, 8.- 10.7.1995" (CF); 26 4,1%, "CINA - Qinghai m. 4400-4600, from Little Maduo to the south sour- ce of Yellow River, 14.VIII.1988" (CS). DERIVATIO NOMINIS. Dedicated to my friend Dominique Echaroux from Dourdan, France. DESCRIPTION. Length 4.26 to 4.48 mm. Piceous black with strong bronze reflections (gree- nish in the paratypes from Sichuan). Antennae piceous black (antennomere 1 red in the paratypes from Sichuan). Legs dark brown to piceous black. Head as in 5. baehri but with deeper frontal furrows. Antennomeres rather short and eyes flat, less than in B. baehri. Head and pronotum in some specimens rugose. Pronotum (pw/pl= 1.30 to 1.36). Basal foveae superficial. Hind rectilinear part of the side, before hind angles, shorter than in the other species. Median impression not strongly impressed. Hind angles square, without lateral carina. Basal transverse impression rather superficial. Base rugose and basal foveae rather superficial. Legs short. Elytra (el/ew = 1.45 to 1.46) strongly ovate with rounded shoulders and maximum width at the hind third. Apex strongly rounded. Elytral striae 1 to 7 superficial, punctate. Basal margin reaching a point between the beginning of stria 5 and 6; apical stria strongly reduced, almost absent. Intervals flat. Microsculpture in isodiametric meshes, coarse on the neck, superficial on the front in few specimens, very superficial at the sides of the pro- notum and evident on elytra. Surface matt alutaceous. Male genitalia (fig. 61). Median lobe of aedeagus wider than in B. baehri, with long main sclerite, ribbon brush strongly sclerotized, and large basal chitin platelet flat, tran- slucent, with strongly cheratinized edges in the ventral part. Basal part of the main scleri- te and base of the other mentioned sclerites slightly protruding from the basal opening. Ventral border of the median lobe strongly sclerotized, dark. Left paramere with four setae, right one with three. Female genitalia (fig. 63). Reservoir of spermatheca "bean shaped", apparently divi- ded in two cavities. DISTRIBUTION (fig. 92). China: Tibet, Sichuan, Qinghai. AFFINITIES. See under B. maddisoni n. sp. B. (Ocydromus) maddisoni n. sp. (fig. 52) DIAGNOSIS. An Ocydromus of the baehri group, relatively large with elytra rather elonga- te but with maximum width clearly behind the middle, and wide and large median lobe of the aedeagus. TYPE LOCALITY. China - N Tibet, Tangula Shan Mts., Tangula Pass, 5100-5300 m. 48 TOLEDANO TYPE SERIES. Holotype, d, "China - N Tibet, Tangula Shan Mts., Tangula Pass, 5100-5300 m, 8.- 11.7.1997" (CT); 96 4,798, paratypes, same date and locality as the holotype (CT). DERIVATIO NOMINIS. Dedicated to my friend Dr. David Maddison, father of the Tree of Life Project, well known bembidiologist and kind reviser of my poor English language. DESCRIPTION. Length 4.24 to 4.40 mm. Piceous - black metallic with greenish reflections. Legs red-brown. Antennomere 1, base of antennomeres 2 and 3 and apex of antennomere 11 brown, rest of antennae piceous. Head glossy with very superficial frontal furrows. Few very superficial punctures behind the frontal furrows. Eyes convex as in B. echarouxi. Antennomeres slightly more elongate than in B. echarouxi. Pronotum transverse (pw/pl = 1.31 to 1.40), glossy, with basal foveae triangular, dee- per than in B. echarouxi, median line slightly impressed, basal transverse impression deep, base finely punctate. | Legs rather short. Elytra (el/ew = 1.50 to 1.53) rather elongate, with maximum width behind middle. Humera rather narrow, rounded, sides of elytra slightly diverging posteriorly, ending in an extremely rounded apex. Striae 1 to 7 evident but rather superficial, ending slightly befo- re apex, intervals flat. Elytra rather shiny, even in the specimens with coarse elytral micro- sculpture. Microsculpture isodiametric superficial on the neck, very superficial on the sides of the pronotum, and evident on the whole elytra. Male genitalia (fig. 62). Similar to that of B. echarouxi, but with ribbon brush and basal chitin platelet longer, more extended in the direction from the base to apex of the median lobe. Ventral margin of the median lobe less sclerotized than in B. echarouxi. Left paramere with four setae, right one with three. DISTRIBUTION (fig. 92). China: Tibet. AFFINITIES. Surely stictly related to B. echarouxi n. sp. as indicated by the structure of the median lobe of the aedeagus, almost identical, and the general body shape. Bembidion (Ocydromus) gr. radians DIAGNOSIS. Group of oriental species within the subg. Ocydromus sensu Kryzhanovskij et al. (1995) characterized by relatively small size (3.60 to 4.72 mm), the metallic integument of head, pronotum and elytra and the absence of a long main sclerite in the median lobe of the edeagus. SYSTEMATIC NOTES. Andrewes (1935) grouped few species in the radians group. As men- tioned above, the species groups of Andrewes do not have systematic value. However, some of the species of this group actually are related to each other. Thanks to the courtesy of Dr. Hine, of the Natural History Museum, London, I could study the types of most species of the radians group. In my opinion B. radians and B. charon can be attributed to the subg. Ocydromus. While I could observe that B. endymion Andrewes, 1935 surely belongs to Peryphophila Netolitzky, 1942 (see below), the remaining species (B. cymindulum Andrewes, 1930, B. psuchrum Andrewes, 1922 and B. algidum Andrewes, 1935) need further inve- stigation in order to discover their correct systematic position. Systematic notes on the paleartic Bembidion, mainly from China 49 ? Tresa tea a Re :$ è è 3: A. & Ro & ®: $ è #i: è 2) di: è fn à eae ee Figs. 64-69. Photographs of the habitus of: 64 - B. radians, holotype; 65 - B. radians puetzi, holoty- pe; 66 - B. radians shaanxianum, holotype; 67 - B. charon, holotype; 68 - B. moritai, holotype (magni- fication a); 69 - B. yunnanum (magnification b). 50 TOLEDANO KEY TO THE SPECIES OF THE RADIANS GROUP SENSU NOVO. 1 - Evident punctures behind frontal furrows; elytra markedly convex, pronotum rather parallel at SiS na Schaan ee re Jorn I a al leds bilans, Aa kan sl th moritai n. Sp. - Head with or without punctures, elytra less convex and pronotum more sinuate at sides ..... 2 2 - More or less evident punctures behind frontal furrows; pronotum with base slightly larger than the anterior margin, and less convex; elytral striae well punctate; median lobe of the aedeagus OI AES n n) LS OI. 3 - No punctures behind frontal furrows; pronotum with base slightly narrower than the anterior mar- gin and more convex; striae less impressed; median lobe of the aedeagus more curved ventrally eee ivi A o RIE charon Andrewes, 1926 3 - Elytral microsculpture absent or poorly developed; India, China (Shaanxi) ............... + - Elytral microsculpture in fine transverse meshes on the elytra; southwestern China (Sichuan, E (fonica ee, e ae te ss vo RR radians puetzi n. ssp. 4 - Microsculpture sometimes present, not well developed, upper surface dark bronze, sometimes Wwdthrorscenisktrefiecrions; Nindia Lele... radians radians Andrewes, 1922 - Microsculpture always absent, upper surface bronze, with lighter shoulders; China (Shaanxi) . . MS a ol 0. radians shaanxianum n. ssp. B. (Ocydromus) radians ssp. radians Andrewes, 1922 (fig. 64, 70) Bembidion radians Andrewes, 1922 This is one of the smallest species of subgenus Ocydromus. My attribution of the spe- cies to Ocydromus 1s due to the presence of the discal elytral pores almost on stria 3, with basal elytral margin normal, not angulated in respect of the lateral margin, reaching the beginning of stria 5. The character of the punctures behind the frontal furrows is shared by the Ocydromus species group sensu Kryzhanovskij et al. (1995). Nevertheless, this cha- racter, used in the past for subgeneric diagnosis, is a character often recurring within other species groups of subg. Ocydromus (e.g. lenae group, lunatum group) and outside the sub- genus (e.g. B. (Synechostictus?) cameroni Andrewes, 1922, Bembidion (Talanes) asperi- colle Germar, 1829, Bembidion (Princidium) punctulatum Drapiez, 1820) and it does not necessarily indicate relationships. The occurrence of this species in China with two new subspecies is reported below. EXAMINED MATERIAL. Holotype, 4, "Gopaldhara Sikkim, H. Stevens" (BMNH); 16, 1%, Paratypes "Sikkim, Lachung, H. Stevens" (BMNH); 19, North India-West Bengal, Shingalila National Park, Shirikhola, 18.-28.5.1999, 2600m (CT) AFFINITIES. Surely related to B. charon as indicated by the body structure and male geni- talia. B. (Ocydromus) radians puetzi n. ssp. (fig. 65) DIAGNOSIS. A subspecies of B. radians differing from the typical form by the presence of more evident elytral microsculpture in fine transverse meshes and by the more depressed elytra. TYPE LOCALITY. China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan). TYPE SERIES. Holotype, à , "China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 1. 2100 m, 29.36 N / 102.04 E, 27./28./31.V.97" (CW); paratypes, 1 d , same date and locality as the holotype; 1d, "China: W-Sichuan: (13), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Systematic notes on the paleartic Bembidion, mainly from China a Camp 1. 2100 m, 29.36 N / 102.03 E, 27.-31.V.97" (CSk); 48 4,29 2, "China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 2. 2500-2700 m, 29.35 N / 101.02 E, 30./31.V.97" (CW, CT); 18, 32 2, "China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 2>1. 2500-1940 m, 31.V.1997" (CW); 15, "China: W-Sichuan: (13), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 1. 2100 m, 29.36 N / 102.03 E, 27./28./31.V.97" (CW); 1 9, "China: W-Sichuan (5), Daxue Shan, Mu Ge Cuo, NW Kangding, 30.10.57N, 101.52.09E, 3200-3400 m, 21.05.1997" (CSk); 22 2, "China, Sichuan, Daxue Shan, Gongga Shan Mt., Hailougou Glacier Park, env. Glacier Tongue, ca. 3.5 km above Camp. III, ca. 3200 m, 29.V.1997" (CPz); 12, "China, Sichuan, Daxue Shan, Gongga Shan Mt. Hailougou Glacier Park, way from Camp II to Camp I, 2620 m-1940 m, 31.V.1997" (CP); 264 3,49 2, "China, Sichuan, Daxue Shan, Gongga Shan Mt. Hailougou Glacier Park, env. Camp II, 2650 m, sifted, 30.V.1997" (CPz); 1d, "China, Sichuan, Ganzi pref., Daxue Shan, 101.57 E, 30.03 N, N Kangding, 2600-2700 m, 22./14.V.1997" (CPz); 16, 19, "China, Sichuan, Ganzi pref., Daxue Shan, 101.52 E, 30.11 N, Mugecuo ca. 26 km NW Kangding, 3200-3400 m, 21.V.1997" (CPz); 56 8,3% 2, "China / Sichuan, 103.20 el / 29.30 nw, Mt. Emei 2600 m, 4.- 18.V.1989" (NMB, CT); 56 8,72 9, "China, pr. Sichuan, Emei Mt. 2500 m, 4.-20.5.1989" (NMB, CT); 26 & , "W Sichuan, 2-6.VIL.1994, 29.36 N 102.06 E, 1900-2900 m, Gonggashan - Hailuogou" (NMB, CT); 38 8,32 2, "W Sichuan, 3-6.VII.1994, 29.35 N 102.00 E, 2900-3200 m, Gonggashan - Hailuogou" (CF, CT); 1 9, "China - S Sichuan, Daliang Shan mts. 9.-11.VI.1998, road Meigu-Leibo vill., pass 15 km NE of Meigu, 28°25'N, 103°17'E" (CT); 26 9,32 2, "China, Sichuan 7 July 1993, Mt. Emei, 3100 m, 180 km S of Chengdu" (CT); 1 £ "Cina - W Sichuan, Emei Shan, VII. 1994" (CS); 236 (*) "China / Sichuan, 103.20 el / 29.30 nw, Mt. Emei 2600 m, 4.-18.V.1989" (NMB, CT); 29 © (*) "China, pr. Sichuan, Emei Mt. 2500 m, 4.-20.5.1989" (NMB); 19 (*) "Cina - Sichuan, Bangu - Emei Dh., VII 94 (CS); 88 4,28, "China S. Sichuan (Ya' an Pref., Shimian Co.) Xiaoxiang Ling pass betw. Shimian Ganluo, 27 km SE Shimian, 29°03N/102°31E, 2450 m, 8.VII.1999" (CW, CT); 1g, "China W Sichuan (Ganzi Tibet Aut. Pref. Luding Co.), W Erlang Shan Pass, 2600m, 7 km SSE Luding, 29°51N/102°15E, 20-29.VI.1999" (CW); 126 4,29 2, "CHINA: S-Sichuan Ya’an Prefecture, Shimian Co. Xiaoxiang Ling, Pass zw. Shimian u. Ganluo, 27 km SE Shimian, 29°03N 102°31E, 2450m, Quellsumpf, Bachufer, 8. VII. 1999" (CW); 22 2, "CHINA: W-Sichuan Ya’an Prefecture, Tianquan Co., W Erlang Shan Pass, 2900m, 29.VI.1999, 29.51.13N, 102.17.28E, sifted" (CPz); 19, "CHINA: W-Sichuan Aba Tibetan Aut. Pref., Weizhou Co., Qionglai Shan, Wolong valley, 69 km WSW Guanxian, 3500m, 30.53.57N, 102.58.63E, 15.VII.1999, mix forest sifting" (CPz); 16, "CHINA: W-Sichuan Ganzi Tibet. Aut. Pref., Kangding Co. Daxue Shan, Mu Ge Cuo, 2 km oberhalb unt. See, 30°11N, 101,52E, Bachufer 5. VII. 1999" (CW); 16, "China Quinghai reg. 3500 m, 120 km W of Qinghai Hu, Tianjun 3.-4.7.1990" (CP); 14, "China. Gansu, 70 km Wudu, 1800-2600 m, 2.6.'97" (CF); 22 9, "China, Gansu, Win Shan range, 70 km N.W. Wudu, 1-VI-97" (CBr); 19, "C China W- Sichuan, Luding co., Moxi env., VI 1993, Gongga Shan, 3000 m" (CB1); 34 d, 19, "China: Sichuan, Gongga Shan, Hailogou, above Camp 3, 29°35N 102°00E, 2800-3300m, 6-8.VII.98" (CMb); 26 d "CHINA: Sichuan 103.20e1/29.30nw Mt. Emei 2600 m 4.-18.V.1989, S.&J. Kolibac" (NMB, CSc). The specimens marked with (*) have square shoulders. DERIVATIO NOMINIS. Dedicated to my friend Andreas Piitz, in whose collection I found some paratypes of this subspecies. DESCRIPTION. Length 3.68 to 4.28 mm. Metallic dark green, more rarely bronze, legs red- dish with knees slightly darkened, antennomere 1 and base of 2, 3, 4 red, rest of antennae piceous. Head with rather superficial and parallel frontal furrows, not extending to the cly- peus; behind them, a few small punctures. Eyes rather convex and antennae rather short. Pronotum (pw/pl = 1.32 to 1.36) cordate, with base narrower than the anterior mar- 52 TOLEDANO gin. Front angles very slightly more advanced than the anterior margin, rectilinear. Sides strongly curved in the anterior 3/4, then strongly sinuate, with hind fourth, almost rectili- near, forming with the rectilinear basal margin sharp, square hind angles. Front transverse impression evident, median line rather strongly impressed, not reaching the basal margin, basal transverse impression deep, with base coarsely punctate and basal foveae deep, squa- re, punctured, with a laterobasal carina strongly developed. Metasternal process distinctly bordered. Legs relatively long. Elytra ovate (el/ew = 1.45 to 1.49) with rounded shoulders (square in few specimens from Emei Shan). Basal margin reaching the beginning of stria 5. Striae coarsely puncta- te, more superficial toward apex, but normally reaching apex. Only stria 7, more superfi- cial than the others, ending clearly before apex. Scutellar stria long, punctate; apical stria deeply impressed, with apical pore at the middle. | Very superficial microsculpture isodiametric on the neck and in large, slightly tran- sverse meshes on the pronotal sides in some specimens. Elytra with microsculpture in very fine, transverse meshes on the hind half of elytra. Male genitalia (fig. 71). Identical to the typical form. Parameres with three setae. Female genitalia (fig. 82). Reservoir of spermatheca "shoe-shaped", apparently divi- ded into two cavities. A faintly sclerotized "bell-shaped" annulus receptaculi. DISTRIBUTION (fig. 92). China: Sichuan, Gansu. SYSTEMATIC NOTES. Some specimens of the ssp. puetzi from Emei Shan have more deve- loped shoulders. The intraspecific variability of the elytral shape is very high, as in B.csikii Jedlicka, 1937 (Toledano & Sciaky, 1998); curiously this great variability is evident in spe- cimens collected on the same mountain as 5. csikii. The structure of the male genitalia of these specimens is identical to those of the typical form. B. (Ocydromus) radians shaanxianum n. ssp. (fig. 66) DIAGNOSIS. A chinese subspecies of B. radians differing from the ssp. puetzi by the absen- ce of elytral microsculpture, and bronze colour with lighter shoulders, and from the typi- cal form by the shoulders paler in colour. TYPE LOCALITY. China - Shaanxi prov., Qing Ling Shan mts., track Hou Zen Zi vill. to Taibai Shan Mt. 2500 m. TYPE SERIES. Holotype, 6, "China - Shaanxi prov., Qing Ling Shan mts., track Hou Zen Zi vill. to Taibai Shan Mt. 2500 m, 27.-29.VI.1998, mixed forest" (CT); paratypes: 14,39 2, same date and locality as the holotype; 19, "China - Shaanxi, 21-23.VI.1998, Qing Ling Shan mts., road Baoji - Taibai vill., pass 35 km S of Baoji" (CT); 36 4,498, "China (Shaanxi), Qin Ling Shan 108.47 E, 33.51 N, Mt. W pass autoroute km 70, 47 km S Xian 2500-2600 m, 26-29.VIII.1995" (CW). DERIVATIO NOMINIS. Derived from the chinese province where the type material was col- lected. DISTRIBUTION (fig. 92). China, Shaanxi. Known only from the type locality. AFFINITIES. Compared with the ssp. radians this subspecies has the elytra slightly more depressed, as in the ssp. puetzi, and with lighter humera; against ssp. puetzi and a few spe- cimens of the typical form it lacks elytral microsculpture. The elytra of B. radians shaanxia- num are slightly more elongate and parallel than in the two precedingsubspecies. Even thou- gh the taxonomical difference is not too evident, the great geographic isolation from the other form normally without microsculpture, the nominotypical form, led me to describe Systematic notes on the paleartic Bembidion, mainly from China 53 ir Ev PR S Figs. 70-80. Left view of the median lobe of the aedeagus of: 70 - B. radians, paratype, prep.n. 1894; 71 - B. radians puetzi, holotype; 72 - B. charon, hololectotype; 73 - B. moritai, holotype; 74 - B. yun- nanum; 75 - B. niedli, holotype; 76 - B. merum; 77 - B. cnemidotum; 78 - B. misellum, 79 - B. muel- lermotzfeldi, holotype; 80 - B. roberti, paratype. Fig. 81: ventral view of the median lobe of the aedea- gus of B. roberti, holotype. Figs 82-85: reservoir of the spermatheca of: 82 - B. radians puetzi; 83 - B. moritai; 84 - B. roberti; 85 - B. muellermotzfeldi. 54 TOLEDANO this population as a separate subspecies. It is geographically separated from the nomi- notypical form by the ssp. puetzi, clearly distinguishable by the well developed elytral microsculpture, a character rather important for the separation of populations. The male genitalia of B. radians shaanxianum are identical to those of the typical form. B. (Ocydromus) charon Andrewes, 1926 (fig. 67, 72) Bembidion charon Andrewes, 1926 This species also belongs to the radians group sensu novo. In this species, closely related to B. radians as indicated by the habitus and the male genitalia, the punctures on the head are absent: this is another demonstration that is almost impossible to find clear subgeneric divisions among the species of the Ocydromus s.1. EXAMINED MATERIAL. Hololectotype, d, "Gori R. Gorge, N. Kumaon, India, 5-9000 ft. H. G. C." (BMNH), 1 9, Paralectotype, "Gori R. Gorge, N. Kumaon, India, 5-9000 ft. H. G. C." (BMNH), 19, Paralectotype, "Burphu, Gori V., 11500 ft., India, H. G. C." (BMNH), 34 à, 19, India bor. Uttar Pradesh bor., Joshimath, Auli, 2800m, 13-17.7.1994 (CT); 14,1%, India bor. Uttar Pradesh, Badarinath, 4000m, 7.7.1994 (CT); 16, 12, Nepal, Ghorepani, 3000, Pokhara, X.1971, Morvan (CMk, CT) B. (Ocydromus) moritai n. sp. (fig. 68) DIAGNOSIS. An Ocydromus species of the radians group sensu novo, easy to recognize by the markedly convex elytra and the pronotum with sides not too sinuated, and rather squa- te. TYPE LOCALITY. China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan). TYPE SERIES. Holotype, à , "China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 1. 2100 m, 29.36 N / 102.04 E, 27./28./31.V.97" (CW); 106 8,5? 2, same date and locality as the holotype (CW, CT); 14,1%, "China (W Sichuan), Daxue Shan, Hailuogou Glacier Park (Gongga Shan) Camp 2. 2500-2700 m, 29.35 N / 102.02 E, 30./31.V.97" (CW); 14,399, "China, Sichuan, Daxue Shan, Gongga Shan, Mt. Hailougou glacier park, 102.04 E, 29.36 N, river valley ca. 1 km above camp I, 2100 m, 28./31.V.1997" (CPz); 19, "China, Sichuan prov., 27.VI. - 3. VII. 1991, Liziping env., near Shimian, 200 km SW of Ya'an" (CP); 12, "C China S-Sichuan, Emei Shan mt 500-800 m, Baoguo env. VI-1993" (CBI). DERIVATIO NOMINIS. Dedicated to my friend Seiji Morita of Tokyo, Japan. DESCRIPTION. Length 4.12 to 4.72 mm. Upper surface metallic green, legs reddish with femura infuscated. Antennomere 1, 2 and base of the antennomeres 3 to 11 reddish brown; rest of antennae darker. Palps red, only penultimate segment of the maxillary palps sligh- tly darker. Head with normally convex eyes, with some punctures on the front, between the fron- tal sulci, behind the eyes. Pronotum (pw/pl = 1.21 to 1.24) less sinuate than in other species of the group. Hind angles with well developed lateral carina. Base abundantly and coarsely punctate, median line rather deep, reaching the basal border. Metasternal process with a very narrow border. Elytra short (el/ew = 1.46 to 1.48) (ew/pw = 1.62 to 1.67) with marked shoulders, extremely convex. Striae 1 to 7 coarsely punctate, disappearing near the apical fifth. Apical Systematic notes on the paleartic Bembidion, mainly from China 5 stria superficial. Basal border reaching the beginning of stria 5. Discal elytral pores in the third interval, very near stria 3, the anterior at the anterior third, the posterior at the hind third. Microsculpture absent on pronotum and elytra. Male genitalia. (fig. 73). Median lobe of the aedeagus with base markedly angulate with respect to the body. Apex narrow, smoothed, slightly arcuate dorsally. Parameres with three setae. Female genitalia (fig. 83). "Pear-shaped" reservoir of spermatheca, with sclerotized, cylindrical annulus receptaculi. DISTRIBUTION (fig. 92). China, Sichuan. Bembidion (Ocydromus) gr. modestum B. (Ocydromus) yunnanum Andrewes, 1923 (fig. 69) B. niedli Jedlicka, 1965 n.syn. SYSTEMATIC NOTES. The comparison of the types of B. (Ocydromus) yunnanum Andrewes, 1923 and B. (Ocydromus) niedli Jedlicka, 1965 reveals that they are synonyms. In the holotype of B. niedli (median lobe of the aedeagus in fig. 75) the apex of the median lobe of the aedeagus is only slightly more elongate than in the rest of the examined specimens (fig. 74), but this is surely an individual aberration. The diagnostic character used by Jedlicka (1965) does not work: the pronotum, described by Jedlicka (1965) as microsculptured in B. niedli and without microsculpture in B. yunnanum, actually lacks microsculpture in all specimens I could see. Therefore: B. niedli Jedlicka, 1965 = B. yunnanum Andrewes, 1923 n. syn. Probably the mistake was due to the fact that the few specimens I could see in Coll. Jedlicka (NMP) labelled as B. yunnanum had a very dirty pronotum. This species shows a variable development of the apical elytral spots. In the typical form they are very well developed, as shown in fig. 69. Some specimens from Sichuan I could examine completely lack the apical spots, being completely dark, or have very faint, almost invisible apical spots. The examination of their male genitalia revealed they belon- ging to B. yunnanum. EXAMINED MATERIAL. 16 , paratype of B. yunnanum:, "Yunnan Fou" (BMNH); 1%, same locality as the paratype (BMNH); 1 d, same locality as the paratype, "Det. Andrewes" (NMP); 1 9, China merid. provincia Yunnan "det. Jedlicka" (NMP); 14 , Holotype of B. niedli, "Kunmin, China 60, let. Kopek" (NMP); 16, 5 9 2, paratypes of B. niedli, same date and locality as the holotype, (NMP, CP); 46 à, 11 9 2, China, Yunnan prov., Heishui 35 km N Lijang, 27.13N, 100.19E, 2.6.-VI.-93 (CE CT); 126 6, 69 ®, China, Yunnan prov., Heishui 35 km N Lijang, 27.13N, 100.19E, 1-19.VII.1992 (CS, CT, NMW); 36 d, 79 2, China, Yunnan prov., Heishui 35 km N Liyang, 27.13N, 100.19E, 18.6-4.7-93 (CT); 108 4,998, China Yunnan, Lugu Lake - Luo Shui, 27.45N, 100.45E, 8.-9.VII.1992 (CBI, CF, CS, CT); 38 gd, 19, China Yunnan, Ninglang env., 27.19N, 100.55E, 6-10.VII—.1992 (CS, CT); 26 6,29 ®, China: NW Yunnan, Yulongxueshan NP, near Baishui, ca. 30 km N Lijang, 2800-3200 m, 7-11.7.1994 (NMW, CT); 26 4,38, China: NW Yunnan, ca 100 km NW Lijang, Hengduanshan, 56 TOLEDANO Jiduan - Weixi, Jiduan - Ludle, 2200 m, 1.7.1994 (NMW, CT); 26 4,29 9, China, Yunnan, Z Xiaguan, 12 km s Weishan, Weibaoshan, 2500-3000 m, 1-17.7.1993 (NMW, CT); 128 4,59 9, China - Yunnan prov., 22-27 July 1998, Dali old tower env. (CT); 19, China - Yunnan prov., Dali W env., 19.VI.1997 (CT); 26 d, China, Kangding m. 2600, Sichuan, 8.VII.1992 (CS); 1d, Cina - W Sichuan, Kangding 2600 m, 12.VII.92 (CS); 26 4,29 ©, China - S Sichuan, 27°45' N, 101°13' E, pass 20 km S Muli/Bowa, mixed forest, ca 3500 m, 25.7.1995 (CT); 35 4,1%, China - S Sichuan, 27°55' N, 101°19' E, 15 km NW Muli (Bowa), mixed forest, cca 3100 m, 30.VI.1998 (CT); 54 6, 49 2 (*) China - S. Sichuan, Daliang Shan mts., 27. VII. 1997, road Meigu - Leibo, pass 15 Km NE of Meigu, 28°25'N, 103.17'E (CT); 16, 12 (*) China - S. Sichuan, Daliang Shan mts., 9.-11. VI. 1998, road Meigu - Leibo, pass 15 Km NE of Meigu, 28°25'N, 103.17'E (CT); 36 4,49 £ (*) China S. Sichuan, Daliang Shan mts., Zhaojue vill. env. pass Xichang - Meigu vill. 12.-14.VI.1998 (CT); 36, 19 (*) China, S. Sichuan, 7.VII.1998, 27.38N, 102.48E, 10 km SW Butuo, cultural steppe (CT); 56 8,6% 2, "China S. Sichuan (Ya' an Pref., Shimian Co.) Xiaoxiang Ling pass betw. Shimian Ganluo, 27 km SE Shimian, 29°03N/102°31E, 2450 m, 8.VII.1999" (CW, CT); 14 (*) China - S Sichuan, road Xichang - Yanyuan, pass 15km SW Pingchuan, 27.33N, 101.49E, cca 3200 m, 26-27.VI.1998 (CT); the specimens marked with (*) are completely dark. Bembidion (Ocydromus) gr. decorum B. (Ocydromus) hasurada, Andrewes, 1924 (fig. 86, 87) B. merum, Jedlicka, 1933 B. kinfushanum Jedlicka, 1958 n.syn. SYSTEMATIC NOTES. The chinese B. merum Jedlicka, 1933 (fig. 86, median lobe of the aedea- gus in fig. 76) has been synonymized by Miiller Motzfeld (1986b) with the indian B. hasu- rada Andrewes, 1924. In my opinion also B. kinfushanum Jedlicka, 1958 (fig. 87), from southwestern China, is also a synonym of B. hasurada. The type of B. kinfushanum seems only be a small specimen of B. merum with reduced punctuation behind the frontal furrows and with slightly more rounded humera. These characters, together with the more rounded shoulders and the pronotum slightly more depressed, are the diagnostic characters used by Jedlicka (1965) to separate B. kinfushanum from B. merum. I could examine a large series of specimens with many transitional forms between the two Jedlicka taxa. Therefore: B. kinfushanum Jedlicka, 1958 = B. hasurada Andrewes, 1924 n. syn. A comparison of indian and chinese specimens of B. hasurada reveals that chinese ones more frequently have the shoulders better developed and the dimensions are slightly lar- ger on average. The placement of B. hasurada in the decorum group has been noted by Miiller- Motzfeld (1986). EXAMINED MATERIAL. 1 9, N India Kashmir, road to Ladakh, Sonamarg, 6/1995 (CT); 76 6,99 6, N India, Himachal Pradesh, Kulu Valley - Manali, 5/1995 (CT); 13, Nepal, V.1980, Jumla, Rara Lac, 3000 m (CT); 12, Holotype of B. merum:, "Tatsienlu, Prov. Sichuan, China Merid." (NMP); 19, paratype of B. merum:, same locality as the holotype (NMP); 19, Holotype of B. kinfushanum, Systematic notes on the paleartic Bembidion, mainly from China DI "Szechuan mer., Mts. Kinfushan, 2000 m pr. flum. Sung-Kahno" (NMP); 136 6,4 ?, China Sichuan, Ganzi pref., Daxue Shan, 101.57 E, 30.33 N, N Kangding, 2600-2700 m, 22/24.V.1997 (CPz, CT); 365,32 2, Cina - W Sichuan, Kangding 2600 m, 12.VII.1992 (CS, CT); 14, Cina - W Sichuan, Mugezo L., 4000 m, 16.VII.92 (CS); 78 d, 69 2, China (W Sichuan), Daxue Shan, W env. Kanding, 2600-2700 m, 30.03N / 101.57E, 22/24.V.1997 (CW, CT); 16, China: W Sichuan (7) Daxue Shan, W Kanding, 30.03.13N, 101.57.44E, 2700-2800 m, 24.05.1997 (CSk); 1 China (W Sichuan), Daxue Shan, river valley 5 km E Kanding 2500-2800, 30.03 N / 102.00 E, 20/23.V.1997 (CW): 16, China 7.92, Sichuan prov., Kangding (CF); 19, China: Sichuan (18) Qingcheng-Shan, NW Chengdu, 600 m, Flussufer, 30.55N, 103.30E, 4.06.1997 (CSk); 1d, 12, China (C Sichuan), Qincheng Shan, NW Chengdu, 600 m (river bank) 33.55 N / 103.30 E, 3/4.VI.1997 (CW); 38 8, 12, China, Sichuan, 22.6.1993, Dayı Dafeishui Forest, cca 110 km W of Chengdu (CT); 14 , China Sichuan pr., Liziping, 28.6-3.7.1991 (CW); 12 China - S Sichuan, Daliang Shan mts., road Meigu-Leibo vill., pass 15 km NE of Meigu, 28°26'N, 103°17'E, 9-11.VI.1998 (CT); 16, 12, China (Shaanxi), Qin Ling Shan 110.05 E, 34/27 N, Hue Shan, 118 km E Xian, N valley 1200-1400 m, leafy wd., 18/20.VIII.1995 (CW, CT); 76 8,4% 2, China W Sichuan (Ya'an Pref. Tianquan Co.) Jiajin Shan, valley below Labahe N. R. St., 54 km W Ya' an, 30°03N/102°27E (brook cleft) 1500 m, 12.VII.1999 (CW, CT); 16, China: S-Sichuan, Ya'an Prefecture, Shimian Co., Xiaoxiang Ling, Pass zw. Shimian u. Ganluo, 27 km SE Shimian, 29°03N, 102°31E, 2450m, quellsumpf, Bachufer, 8. VII. 1999 (CW); 19, China - Sichuan prov.; Kangding distr. 7.92, Mugeguo lake 4500m (CW). Bembidion (Ocydromus) gr. cnemidotum SYSTEMATIC NOTES. A japanese species, B. (Ocydromus) cnemidotum Bates, 1883 (fig. 88), present also on the Sakalin peninsula and in the Ussuri region, is kept in its own group of species of Ocydromus by Kryzhanovski et al. (1995). The pronotal structure of B. cnemi- dotum is extremely similar to that of B. yunnanum Andrewes, 1923, but the structure of the inner sac of the aedeagus is completely different. I have examined a series of specimens of the japanese B. misellum Harold, 1877 (fig. 89). This species shares the body structure of B. cnemidotum, except for the size, much larger in the latter, and for the structure of the apical pore, isolated in B. misellum and on the apical stria in B. cnemidotum. But the struc- ture of the inner sac of the aedeagus of both species (figs. 77, 78) is strikingly similar. Therefore I include here B. misellum in the cnemidotum group. Morita (1994a) showed the similarities in habitus and in male genitalia between B. cnemidotum and the japanese B. kamikochii Jedlicka, 1965. In the description of the japanese B. ohkurai Morita, 1992, the author compared the new species with B. misellum because of similar habitus and male genitalia. In my opinion B. cnemidotum, B. misellum, B. kamikochii and B. ohkurai belong to the same species group, even though I cannot confirm the placement of the latter since at present I could not examine any specimen belonging to this taxon. EXAMINED MATERIAL. B. misellum: 38 8,3% 2, Japan, Marunuma, Gunma pref., 22-VI-1982, S. Morita leg. (CM, CT); 48 8,4% 2, Japan, Hasuike, Nagano pref., 9.6.74 (CE, CT). B. cnemidotum: 18 8, 12 2, Japan, Oigawa River, Shizuoka, 25.4.71 (CE, CT); 1088,72, S.Sakhalin, Novoalexeevka vill. env., 25-30.05.98 (CT); 16, Rossia or. Sakhalin, Krijon, Kalinino env., 4-9.7.1992 (CT). B. kamikochii: Holotype, 2, "Kamikochi, Japan Alps, 21.VII.1951" (NMP). 58 TOLEDANO Bembidion (Ocydromus) gr. terminale (Bembidion subg. Terminophanes Müller-Motzfeld, 1998) SYSTEMATIC NOTES. The species of the group of B. terminale Heer, 1841 have been inclu- ded by Müller-Motzfeld (1998) in a new subgenus, Terminophanes Miiller-Motzfeld, 1998. The characters used by the author to separate the Terminophanes species are the absence of punctuation at the pronotal base and, mainly, the presence of a particular sclerite ("tri- corned body", Lindroth 1965) in the median lobe of the aedeagus. Even though the species of this group share strict systematic relationships to each other, I prefer to place them into a species group of Ocydromus and attribute, at least provisionally, the new species descri- bed here to the subg. Ocydromus. B. (Ocydromus) roberti n. sp. (figs. 90) DIAGNOSIS. Very elongate species of the terminale group showing flat elytra with shoul- ders at an obtuse angle, maximum elytral width behind the middle, very long temples, redu- ced eyes and extremely elongate antennae. The lateral carinae of the pronotal hind angles are present, even though only suggested by a poorly developed process. In the median lobe of the aedeagus, the lateral right position of the ostium, a charac- ter shared with some other species of the group, is here evident for the very large ostium, extended in its apical half. TYPE LOCALITY. China, Qinghai, Dulan. TYPE SERIES. Holotype, à , "China - Quinghai, Dulan, 10.7.1990" (CS); Paratypes: 26 6,29 9, same data and locality of the Holotype (CS, CT); 1d, "China, Qinghai, Laji Shan range, Chan-Hu villg., VII-97" (CBr); 26 8,32 9, "China, Gansu reg. 4200 m, Dogcanglhamo, 12-15.7.1990" (CP, CT). DERIVATIO NOMINIS. I dedicate this species to my beloved father, Dr. Roberto Toledano, great dentist and astronomer who taught me dentistry and life, by his example, on the occa- sion of his 76th birthday. DESCRIPTION. Length 4.20 to 4.56 mm. Head and pronotum dark brown to piceous, elytra reddish - brown to piceous, with faint metallic reflections; in the specimens from Gansu (fig. 95) head, pronotum and elytra darker, piceous - black with greenish metallic reflec- tions; legs red; Antennomere 1 to 3 and base of 4 red, rest of antennae dark brown. Head with flat and rather small eyes, temples long, antennae elongate, with anten- nomere 3 and 4 extremely developed, length almost two times antennomere 2. Frontal fur- rows parallel, uneven, rather deep, reaching the half of the clypeus, rarely exceeding poste- riorly the hind third of the eyes. Pronotum narrow (pw/pl= 1.24 to 1.28) cordate, depressed, with sides strongly sinua- te before the hind angles, and with basal margin shorter than the anterior one. Hind angles sharp, laterally prominent. Basal foveae not deep. Lateral carina rudimentary. Base of pro- notum rectilinear at middle, very slightly oblique at sides. Base of pronotum about one half of base of elytra. Lateral seta at the anterior 2/5, posterior seta at the hind angle. Metasternal process rebordered; legs elongate. Elytra (el/ew = 1.56 to 1.58) (ew/pw = 1.73 to 1.76) with square shoulders, sides almost rectilinear, posteriorly diverging to about the hind 3/5, where the width is maximal, Systematic notes on the paleartic Bembidion, mainly from China 39 and then converging, rounded, to the apex. Basal elytral margin ending at beginning of stria 5. Elytral striae punctate, normally impressed on the disk, more superficial at sides, almo- st disappearing after the posterior 3/5, only stria 1, rarely stria 2 reaching the apex. Apical stria short, not too deep but evident. Stria 7 absent. Posterior discal elytral pore on stria 3, at the posterior 3/4 of elytra. Anterior discal pore on stria 3 at the anterior 2/5 of elytra, visible only in the specimens from Gansu. Microsculpture of elytra and pronotum with iso- diametric meshes, extended on the whole surface but superficial, so that the integument is rather shining. Male genitalia (figs. 80, 81). Ostium of the median lobe of the pronotum very large, opened on the right side of the apex, from the apical extremity, backward to the half of the length. On the right side, about at the anterior third of the median lobe, a little bridge of cheratinized tissue links the ventral with the dorsal margin. Tricorned body strongly deve- loped, its dorsal part dorsally surrounding the apical part of main sclerite, basal end of main sclerite and ribbon brush slightly protruding from the basal opening of the median lobe. Left and right paramere with three setae. Female genitalia (fig. 84). Spermatheca simple, with reservoir small, "bean shaped", with one main lobe cylindrical and another one, smaller, length one half of the main one, slightly bent with it. A sclerotized annulus receptaculi is not present. DISTRIBUTION (fig. 92). China: Qinghai, Gansu. AFFINITIES. The structure of the pronotum of B. roberti n. sp. is similar to the typical one of B. (Ocydromus) of the terminale group, even though in B. roberti the pronotum is nar- rower than in other species of the group. Because of the extreme development of the ostium of the median lobe of the aedeagus, I initially planned to describe B. roberti as belonging to anew subgenus. I prefer to give to this shape of ostium only a specific value, to prevent any unnecessary further separation within the Bembidiini, even though a relatively inde- pendent position of B. roberti within the terminale group is rather evident also by the habi- tus of the species. B. (Ocydromus) muellermotzfeldi n. sp. (fig. 91) DIAGNOSIS. A species of the terminale group very similar to B. mckinleyi Fall, 1926 and its subspecies, easy to recognize because of paler legs, elytra slightly more convex and the form of male genitalia. TYPE LOCALITY. China, W Sichuan, 3800 m, road Luhuo-Sertar, 20 km N Luhuo, 31°32'N 100°42'E. TYPE SERIES. Holotype, & , "China, W Sichuan, 3800 m, road Luhuo-Sertar, 20 km N Luhuo, 31°32'N 100°42'E, mixed forest, 21.VII.1997" (CT); paratypes: 29 2, same date and locality as the holotype (CT); 12, "China, W Sichuan, 4000 m, road Luhuo-Sertar, 40 km N Luhuo, 31°41'N 100°44'E, thuya and picea forest, 23.VI.1997" (CT); 76 8, 122 9, "China, W Sichuan, (Ganzi Tibet. Aut. Pref. Kangding Co.), Daxue Shan, brook bank W Tsheto La Pass, 3650 m, 20 km W Kangding, 30°04 N/101°46 E, 25. VI. 1999" (CW, CT); 26 5, 22 2, "CHINA: W-Sichuan Ganzi Tibet. Aut. Pref., Kangding Co. Daxue Shan, Bachufer W Tsheto La Pass, 3650m, 20 km W Kangding, 30°04N, 101°46E, 25. VI. 1999" (CW); 45 6, 82 2, "E Tibet, road Rawu - Baxoi, pass 10 km N Rawu, 29°38'N 96°43'E, 4300 m, alpine meadow, 12.VII.1997" (CT); 54 4,6% 2," E Tibet, Bamda env., 4400 m, 30°15'N, 97°16'E, grassland, 5.VII.1997" (CT); 3¢ d, 39 2, "China Gansu reg. 4200 m, 60 TOLEDANO Dogcanglhamo, 12-15.7.1990 (CP, CS, CT); 26 4,29 9; "China: W Sichuan, 3500-4300m, Temple 35 km Sabdé, 29°40N 101°20E, 13-14.V11.1998" (CMb). DERIVATIO NOMINIS. I dedicate this species to Dr. Prof. Miiller-Motzfeld, great bembidio- logist, who honours me by sending his interesting comments on my works. DESCRIPTION. Length 5.20 to 5.50 mm. Dark metallic green, shiny. Antennomere 1, 2, base of 3, 4 and apex of 11 red to reddish brown. Rest of antennae piceous. Legs red with femurs slightly infuscated. Head rather narrow with frontal furrows deep and very slightly converging, exten- ding to clypeus, where they reach the lateral clypeal setae. Front impunctate. Eyes sligh- tly reduced. Antennae elongate. Pronotum (pw/pl = 1.33 to 1.38) cordiform, with base narrower than the anterior mar- gin, rectilinear, sides rounded, sinuate at the hind fifth, and then slightly diverging, forming with the rectilinear base hind angles that are sharp, acute. Anterior transverse impression shallow. Median line evident, almost reaching basal margin. Basal foveae rather superfi- cial, rugose, without laterobasal carina (present, but extremely rudimentary in few speci- mens). Lateral seta at the anterior 2/5, posterior seta at the hind angle. Legs long and sharp. Metasternal process distinctly bordered. Elytra elongate (el/ew = 1.58 to 1.62) evidently larger than pronotum (ew/pw = 1.64 to 1.66), with maximum width posteriorly to the middle, with square shoulders. Basal mar- gin reaching the beginning of stria 5, striae rather deeply punctate-sulcate, reaching apex, even though more superficial near apex. Stria 7 absent. Scutellar stria punctate, long. Apical stria strongly reduced, superficial and short. Elytral discal pore punctures in interval 3, adjoining stria 3. Microsculpture superficial, isodiametric on neck, very faint, difficult to see, at the pronotal sides in a few specimens; superficial, but evident, in transverse meshes at the ely- tral apex. Male genitalia (fig. 79). Having the typical structure of the group, the "tricorned body" (Lindroth, 1965). The inner sac is slightly protruding from the basal opening of the aedea- gus, as in most species of the terminale group. Compared with that of B. mckinleyi, the median lobe is clearly narrower and without the slight convexity at the ventral margin. Parameres with four setae. Female genitalia (fig. 85). Reservoir of spermatheca almost identical to B. roberti. DISTRIBUTION (fig. 92). SW China (W Sichuan, E Tibet) AFFINITIES. Within the Terminophanes group this species seems to be rather closely related to Bembidion mckinleyi Fall, 1926, from which it can be easily distinguished for the lighter color of the appendages, the less parallel elytra and the differences in the male genitalia. Bembidion subg. Peryphophila Netolitzky, 1942 B. (Peryphophila) endymion Andrewes, 1935 I attribute here B. endymion to the subg. Peryphophila because of general habitus, characterized by large, oval elytra and rather small, depressed pronotum, the unbordered Systematic notes on the paleartic Bembidion, mainly from China 61 Figs. 86-91. Photographs of the habitus of: 86 - B. merum, holotype; 87 - B. kinfushanum, holotype; 88 - B. cnemidotum; 89 - B. misellum; 90 - B. roberti, paratype from Gansu; 91 - B. muellermotz- feldi, holotype. metasternal process and the presence of a slight crista clavicularis at the elytral base. As mentioned above, this is one of the species of the radians group sensu Andrewes (1935). EXAMINED MATERIAL. | 2, Holotype, "Haldwani Dist., Kumaon, India, H.G.C." (BMNH). 62 TOLEDANO Bembidion subg. Pseudolimnaeum Kraatz, 1888 B. (Pseudolimnaeum ) arnosti Nakane, 1978 Bembidion szekessji Jedlicka, 1961 nec Fassati, 1955 Bembidion (Pseudolimnaeum ) jedlickanum Toledano, 1999 n.syn. In my revision of the subg. Bembidion (Toledano, 1999) I have replaced the preoc- cupied name Bembidion szekessji Jedlicka, 1961 with Bembidion jedlickanum Toledano, 1999, but it is an junior synonym of Bembidion arnosti Nakane, 1978 (Werner Marggi, per- sonal communication). Therefore: B. (Pseudolimnaeum) jedlickanum Toledano, 1999 = B. (Pseudolimnaeum ) arnosti Nakane, 1978 n.syn. BIOGEOGRAPHICAL NOTES ON THE CHINESE BEMBIDIINI Many external influences enrich the chinese fauna. Elements of the highlands of Central Asia also live in western China (Xinjang). The fauna of the southwestern China (Yunnan) shows elements in common with the southeastern asian region and northern India. The siberian fauna clearly exerts its influence in Mongolia, where most species of eastern Siberia are present, and also in the chinese provinces of Nei Mongol and Manchuria. Some japanese species are strictly related to some chinese ones, as, for example B. gotoen- se Habu, 1977, strictly related to the chinese B. peleum Jedlicka, 1933 and to B. schoen- manni n. sp.; also the subgenus Aureoplataphus Netolitzky, 1942, considered as mono- specific and endemic to Japan, includes also a chinese species, B. jaechi n. sp. Some rela- tionships between the nepalese and chinese fauna have been shown by the subgenus Hoquedela Müller-Motzfeld, 1988 (Toledano & Sciaky, 1998) and by the genus Amerizus Chaudoir, 1868 (Perrault, 1981, 1985; Deuve, 1998); this last, as discovered by Perrault (1981) reveals some relationships also with the northern american fauna. Nonetheless, the species of Bembidion of southwestern China are in large part endemic, because of extre- me isolation due to the high mountains of this region (Deuve, 1998; Toledano & Sciaky, 1998; Toledano, 1998, 1999). PROVISIONAL CHECKLIST OF THE CHINESE BEMBIDIINI Based on my study of the chinese fauna I include this provisional checklist of spe- cies at present known from China. It is likely that additional taxa now known from neigh- boring regions mentioned in the preceding paragraph may also occur in China. A few species new to China have been identified during the preparation of this work: Bembidion (Notaphus) obliquum Sturm, 1825 EXAMINED MATERIAL. 16 22 9 China Gansu prov., 120 km SW Lanzhou, Ponggartang, 30.VI.- 2.V11.1992 (CS, CT); 1 £ China West Sichuan prov., 60 km S of Hongyuan, ca. 4200 m, 27-29.VI.1991 (CS). Systematic notes on the paleartic Bembidion, mainly from China 63 + ron) ANTO RTE PA URSS TROPIC OF CANCER Fig. 92. Map of China showing the distribution of B. jaechi (1), B. schoenmanni (2), B. baehri (3), B. echarouxi (4), B. maddisoni (5), B. radians puetzi (6), B. radians shaanxianum (7), B. moritai (8), B. roberti (9) and B. muellermotzfeldi (10). Bembidion (Bembidion) quadripustulatum ssp. quadripustulatum Serville, 1821 EXAMINED MATERIAL. | 9, China, Nei Mongol, Wuhai, 8.5.-13.5.1996 (CKu). Bembidion (Plataphus) gebleri ssp. persuasum Netolitzky, 1938 EXAMINED MATERIAL. 26 d, 29 9, NE China: Jilin, 30 km NE Baihe City, nr. Hongai, 17.8.1994, 650 m (15) (NMW, CT). Bembidion (Bembidionetolitzkya) piceocyaneum Solsky, 1874 EXAMINED MATERIAL. 16,1%, China, Xinjiang, 2000-3000 m S Slope of Tien Shan mts., road Kuga - Bayanbulak, ca 100 km NNE Kuga, 8-11.V.1993 (CT); 16, China occ., 16.-21.7.1991, Tian Shan 2000-3000 m, 70 km Urumgi - Hauxia (CP). Tribe Bembidiini gen. Asaphidion Gozis, 1886 championi Andrewes, 1924 cupreum Andrewes, 1925 fragile Andrewes, 1925 granulatum Andrewes, 1925 semilucidum Motschulsky, 1862 64 TOLEDANO gen. Bembidion Latreille, 1802 subg. Chinocillenus Netolitzky; 1942 sinicus Andrewes, 1938 subg. Desarmatocillenus, Netolitzky, 1942 foochowensis Lindroth, 1980 subg. Odontium Leconte, 1848 aeneipes Bates, 1883 gebieni Netolitzky, 1928 subg. Bracteon Bedel, 1879 stenoderum ssp. mukdense Kirschenhofer, 1984 velox Linné, 1761 subg. Eurytrachelus Motschulsky, 1846 pogonoides Bates, 1883 rufotibiellum Fairmaire, 1888 subg. Metallina Motschulsky, 1850 mundatum Netolitzky, 1920 subg. Neja Motschulsky, 1864 gansuense Jedlicka, 1965 subg. Philochtemphanes Netolitzky, 1923 brancuccii n. sp. daliangi n. sp. exquisitum Andrewes, 1923 hansi Jedlicka, 1932 perditum Netolitzky, 1920 = tienmushaniense Kirschenhofer, 1984 n.syn. ? goetzi Jedlicka, 1965 subg. Pekinium Csiki, 1901 (subg. incertae sedis) chinense Csiki, 1901 (species inquirenda) subg. Hoquedela Müller-Motzfeld, 1988 csikii Jedlicka, 1937 subg. Andrewesa Netolitkzy, 1931 incisum Andrewes, 1921 subg. Princidium Motschulsky, 1864 coreanum Jedlicka, 1946 davidi Schuler, 1956 subg. Notaphus Stephens, 1829 obliquum Sturm, 1825 semipunctatum ssp. elegantulum Sahlberg, 1844 subg. Notaphocampa Netolitzky, 1914 niloticum ssp. batesi Putzeys, 1875 subg. Notaphomimus Netolitzky, 1931 sobrinum Boheman, 1848 subg. Microsinocys Toledano, 1998 barkamense Toledano, 1998 daxuense Toledano, 1998 facchinii Toledano, 1998 herbertfranzi Toledano, 1998 Jani Toledano, 1998 Systematic notes on the paleartic Bembidion, mainly from China luhuoense Toledano, 1998 ginghaicum Toledano, 1998 rebeccae Toledano, 1998 schillhammeri Toledano, 1998 schuelkei n. sp. turnai Toledano, 1998 wraseanum Toledano, 1998 subg. Josefia n. subg. belousovi n. sp. luisae n. Sp. pieroi n. sp. shugela n. sp. taguense n. Sp. subg. Bembidion Latreille, 1802 quadrimaculatum ssp. quadrimaculatum Linné, 1761 quadrimaculatum ssp. mandli Netolitzky, 1932 quadripustulatum ssp. quadripustulatum Serville, 1821 sciakyi Toledano, 1999 sciakyi ssp. luguense Toledano, 1999 sciakyi ssp. rinaldi Toledano, 1999 subg. Emphanes Motschulsky, 1850 bulgani Jedlicka, 1968 articulatoides Jedlicka, 1932 gobiense Jedlicka, 1964 subg. Semicampa Netolitzky, 1910 mandarin Netolitzky, 1939 subg. Bembidromus n. subg. panda n. sp. subg. Plataphus Motschulsky, 1864 asiaticum JedliCka, 1965 hamanense Jedlicka, 1933 (subg. Bembidionetolitzkya?) gebleri ssp. persuasum Netolitzky, 1938 speciense Jedlicka, 1932 subg. Parataphus Jedlicka, 1932 heyrovskyi Jedlicka, 1932 subg. Plataphodes Ganglbauer, 1892 farkaci Toledano & Sciaky, 1998 subg. Blepharoplataphus Netolitzky, 1920 davaai Jedlicka, 1968 subg. Trichoplataphus Netolitzky, 1914 hysteron Netolitzky, 1943 kara Andrewes, 1921 lissonotoides Kirschenhofer, 1989 proteron Netolitzky, 1920 tambra Andrewes, 1923 subg. Aureoplataphus Netolitzky, 1942 jaechi n. sp. subg. Bembidionetolitzkya Strand, 1929 piceocyaneum Solsky, 1874 65 66 subg. Ocydromus Clairville, 1806 gr. gr. gr. gr. gr. tetracolum abbreviatum ssp. uvidum Andrewes, 1924 andrewesi Jedlicka, 1932 babaulti Andrewes, 1924 captivorum Netolitzky, 1943 goetzi Jedlicka, 1965 holdhausi Fassati, 1952 insidiosum Solsky, 1874 morawitzi Csiki, 1928 obscurellum ssp. turanicum Csiki, 1928 olemartini Kirschenhofer, 1984 poppii ssp. eugenes Jedlicka, 1933 poppii ssp. pohlai Kirschenhofer, 1984 taiyuanense Kirschenhofer, 1984 lunatum collutum Bates, 1873 infuscatum Dejean, 1831 obenbergeri Lutschnik, 1923 ocylum Jedlicka, 1933 semiferrugineus Kirschenhofer, 1984 semilunium ssp. serorum Netolitzky, 1934 . terminale roberti n. Sp. muellermotzfeldi n. sp. pseudoconsummatum Kirschenhofer, 1984 decorum hasurada Andrewes, 1924 = kinfushanum Jedlicka, 1958 n.syn. = merum Jedlicka, 1933 saxatile saxatile ssp. fuscomaculatum Motschulsky, 1844 modestum scopulinum ssp. thermarum Motschulsky, 1844 = thermarum mongolicum Jedlicka, 1965 yunnanum Andrewes, 1923 = niedli Jedlicka, 1965 n.syn. gr. parepum gr. gr. gr. parepum Jedlicka, 1933 baehri baehri n. sp. echarouxi n. sp. maddisoni n. sp. radians radians puetzi n. ssp. radians shaanxianum n. ssp. moritai n. Sp. lenae chloreum Bates, 1873 TOLEDANO Systematic notes on the paleartic Bembidion, mainly from China 67 peleum JedliCka, 1933 = nanpingense Kirschenhofer, 1984 n.syn. phaedrum Andrewes, 1923 = purkynei Jedlicka, 1932 schoenmanni n. sp. wutaishanense Kirschenhofer, 1984 incertae sedis klapperichi Jedlicka, 1953 spectans Jedlicka, 1933 sterbai Jedlicka, 1965 sjoelanderi Jedlicka, 1965 vaillanti Schuler, 1955 subg. Testediolum Ganglbauer, 1892 gagates Andrewes, 1914 subg. Pamirium Netolitzky, 1920 roborovskii Mikhailov, 1988 ? subg. Synechostictus Motschulsky, 1864 cameroni Andrewes, 1922 exaratum Andrewes, 1924 aeneoviridimicans Netolitzky, 1938 Gen. Amerizus Chaudoir, 1868 subg. Tiruka Andrewes, 1935 barkamensis Deuve, 1998 baxiensis Deuve, 1998 gongga Deuve, 1998 markamensis Deuve, 1998 mourzinei Deuve, 1998 perraulti Deuve, 1998 queinneci Deuve, 1998 songpanensis Deuve, 1998 turnai Deuve, 1998 wrzecionkoi Deuve, 1998 CONCLUSIONS The systematics of Bembidiini is very far from being settled. The main aim of my research is to reorganize knowledge about the eastern palaearctic fauna, and about the whole complex of world Bembidiini. At present, I prefer to keep the species grouped in very few genera, since in my opinion the homogeneity of the tribe is indubitable. This approach to classification of the Bembidiini often does not allow separation of species into small, easy- to-handle subgenera. Two main groups of subgenera of the Bembidiini include most world species: one is that of species more strictly related to the subg. Bembidion s.str. and the other one is that of species more related to the subg. Ocydromus. Some authors, following Perrault (1981) upgraded Ocydromus to genus, but in my opinion this approach does not help, since in a subgeneric division between the entities placed in Kryzhanovskij et al. (1995) as groups of species of Ocydromus is very difficult to find. Moreover, the whole 68 TOLEDANO complex of Bembidion loses its homogeneity, which in my opinion is indubitable, espe- cially in light of a new interpretation of some diagnostic characters as above. Since the systematic division of taxa is always arbitrary, I think that the best thing to do is to use the method that lets us work most easily. This is, in my opinion, the classical one, used by the father of taxonomy of Bembidiini, Dr. Fritz Netolitzky. Using his classical informal divi- sion in species groups, also the main subgenus, Ocydromus Clairville, 1806, huge and in my opinion distributed almost throughout the whole world, can be rather easily studied. In fact, the main problem of the oriental fauna is that several taxa do not belong clearly to the subgenera that in the palaearctic region are very easy to separate from each other. Considering the subgenus Ocydromus as a multiform complex of groups of species, we can rather easily find a correct systematic position for the new taxa related to it, and for many others descri- bed in the past as Bembidion s.l. ACKNOWLEDGEMENTS. I wish to thank Dr David Maddison for the linguistic revision of the text, Dr Josef Jelinek of the Narodni Muzeum Praze, Dr Heinrich Schönmann and Dr Manfred Jäch of the Naturhistorisches Museum, Wien, Dr Martin Baehr of the Zoologische Staatssammlung, Miinchen, Dr Stuart Hine of the British Museum of Natural History, London, Dr Carlo Pesarini and Mr Maurizio Pavesi of the Museo Civico di Storia Naturale di Milano, Mrs Roberta Salmaso and Dr Leonardo Latella of the Museo Civico di Storia Naturale di Verona, Dr Michael Hansen and Mr Ole Martin of the Universitetes Zoologiske Museum, Kgbenhavn and Dr Michel Brancucci of the Naturhistorisches Museum, Basel, for the kind loan of material preserved in their institutions, all the friends that gave me the material of their private collections to study, and my friend Dr Riccardo Sciaky for his constant supervision of my entomological study. As usual, a special thank is due to Rebecca, my invaluable collaborator. 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"Natura Bresciana", Annali del Museo civico di Storia naturale di Brescia, 29: 193-201. SCIAKY R., 1994 - Revision of the Prerostichus subg. Morphohaptoderus and description of ten new species (Coleoptera, Carabidae, Pterostichinae). Koleopterologische Rundschau, Wien, 64: 1- 19; SCIAKY R. & FACCHINI S., 1997 - Microzargus new genus from the mountains of Asia and notes on other Licinini (Coleoptera Carabidae). Bollettino della Società Entomologica Italiana, 129 (1): 51-65. SCIAKY R. & FACCHINI S., 1999 - A review of the Chinese Loricera, with description of a new sub- genus and three new species (Coleoptera Carabidae Loricerinae). In: A. Zamotajlov & R. Sciaky (eds.): Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij, MUISO publ., Krasnodar, pp. 95-108. SERRANO J. & GALIAN J., 1996 (1998) - A review of karyotypic evolution and phylogeny of carabid beetles (Coleoptera). Proceedings of a Symposium (28 August, 1996, Florence, Italy) XX International Congress of Entomology; Museo Regionale di Scienze Naturali, Torino: 191-228 TOLEDANO L. & SCIAKY R., 1997 (1998) - Three subgenera of Bembidion new to China, with descrip- tion of a new species (Coleoptera Carabidae). Bollettino del Museo Civico di Storia Naturale di Venezia, 48: 1-18 TOLEDANO L., 1998 - Microsinocys, anew subgenus of Bembidion Latreille from western and southwe- stern China (Coleoptera: Carabidae). Koleopterologische Rundschau, 68: 27-45. TOLEDANO L., 1999 - Revision of the palaearctic species of the subgenus Bembidion with descrip- tion of three new taxa from China (Coleoptera, Carabidae, Bembidiini). In: A. Zamotajlov & R. Sciaky (eds.): Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij, MUISO publ., Krasnodar, pp. 195-227. VIGNA TAGLIANTI A., 1993 - Checklist delle specie della Fauna d'Italia. Coleoptera Archostemata, Adephaga 1 (Carabidae). Edizioni Calderini, Bologna: 1-51 VIGNA TAGLIANTI A., 1994 (1995) - Un nuovo Ocys dell'Appennino Abruzzese (Coleoptera, Carabidae). Bollettino dell'Associazione Romana di Entomologia, 49 (3-4): 129-136 Indirizzo dell'Autore: L. Toledano, Museo Civico di Storia Naturale, Lungadige Porta Vittoria 9, I-37129 Verona, Italy. e-mail: lucatole@iol.it Mem. Soc. entom. ital., 78 (1): 71-129. 30 aprile 2000 Davide Sassi & Seniz KISMALI The Cryptocephalinae of Turkey, with Informations on their Distribution and Ecology (Coleoptera Chrysomelidae) Abstract - 91 species of Cryptocephalinae of Turkey are listed, on the basis of personal observa- tions and review of literature. For the first time keys are presented for the identification of genera and species from the territory under study. General and local distribution of species, as well biolo- gical notes and synonyms are reported. Cryptocephalus paphlagonius, new to the science, is descri- bed. C. trapezensis Tappes, 1871 is synonymized with C. cribratus Suffrian, 1847; C. bodemeyeri Weise, 1900 is synonymized with C. moehringi Weise, 1884; C. karsantianus Pic, 1914 is synony- mized with C. variceps Weise, 1884. Lectotypes of C. araxidis Weise, 1898; C. bodemeyeri Weise, 1900; C. karsantianus Pic, 1914; C. moehringi Weise, 1884; C. sultani Pic, 1920; C. virens Suffrian, 1847; Pachybrachys misellus (Weise, 1900) and P. pentheri Ganglbauer, 1905 are designated. Neotype of Cryptocephalus trapezensis Tappes, 1871 is designated. C. pelleti Marseul, 1875 and C. volkovitshi Lopatin, 1976 are newly recorded from the country. Male genitalia of the following species are figured for the first time: Cryptocephalus curda Jacobson, 1897; C. moehringi Weise, 1884; C. lederi Weise, 1889; C. peyroni Marseul, 1875; C. sultani Pic, 1920; C. tappesi Marseul, 1868; C. wehnkei Weise, 1882; Pachybrachis laticollis (Suffrian, 1860); P. pentheri Ganglbauer, 1905. Lastly, P mardinensis (Weise, 1900) and P. tessellatus ssp. orientalis (Weise, 1894) are newly recorded from Lebanon. P. mardinensis is also newly recorded from Syria and Israel. Riassunto - Le Criptocefaline di Turchia, con informazioni sulla loro distribuzione ed ecologia. Il lavoro riassume le conoscenze attuali sulla distribuzione delle 91 specie di Coleoptera Chrysomelidae Cryptocephalinae attualmente note per la Turchia. L’elenco delle presenze è redatto sulla base di materiale inedito e su una revisione critica dei dati reperiti in letteratura. Il catalogo riporta la distri- buzione generale e locale delle specie trattate, oltre a note biologiche e sinonimiche. Per la prima volta vengono compilate chiavi di identificazione per il territorio in esame. Viene descritto Cryptocephalus paphlagonius n. sp. C. trapezensis Tappes, 1871 è posto in sinonimia di C. lus cri- bratus Suffrian, 1847; C. bodemeyeri Weise, 1900 è posto in sinonimia di C. moehringi Weise, 1884; C. karsantianus Pic, 1914 è posto in sinonimia di C. variceps Weise, 1884. Sono designati i lectotipi delle seguenti specie: C. araxidis Weise, 1898; C. bodemeyeri Weise, 1900; C. karsan- tianus Pic, 1914; C. moehringi Weise, 1884; C. sultani Pic, 1920; C. virens Suffrian, 1847; Pachybrachys misellus (Weise, 1900) e P. pentheri Ganglbauer, 1905. Viene designato il neotipo di Cryptocephalus trapezensis Tappes, 1871. C. pelleti Marseul, 1875 e C. volkovitshi Lopatin, 1976 sono nuovi per la fauna di Turchia. I genitali maschili delle seguenti specie vengono raffigu- rati per la prima volta: C. curda Jacobson, 1897; C. moehringi Weise, 1884; C. lederi Weise, 1889; C. peyroni Marseul, 1875; C. sultani Pic, 1920; C. tappesi Marseul, 1868; C. wehnkei Weise, 1882; Pachybrachis laticollis (Suffrian, 1860); P pentheri Ganglbauer, 1905. Infine, P mardinensis (Weise, 1900) e P. tessellatus ssp. orientalis (Weise, 1894) sono per la prima volta segnalati per il territo- rio del Libano. P. mardinensis è nuovo anche per la Siria e Israele. Key words: Cryptocephalinae, Turkey, faunistics, new species, synonymy. 72 SASSI & KISMALI INTRODUCTION Cryptocephalinae constitute an important subfamily of Chrysomelidae, distributed throughout the world and distinguished by a combination of the following basic features: body short and oval, small to moderate in size, rounded or obtuse at ends; head is vertical and hidden by the pronotum, eyes are often emarginate on the inner rim, antennae are thin and filiform, pygidium is large and for most part is not covered by elytral surface. Chromatic pattern often includes gaudy colours or metallic iridescence, at least in the genus Cryptocephalus. External features are often very similar in different species and chroma- tic pattern can vary a lot within the same taxon, so the correct identification can require the analysis of male genital characteristics (aedeagus). On the contrary, female genitalia are of little value for taxonomic purposes, according to the present knowledge. Adults are phytophagous and feed on flowers or leaves of a wide range of herbaceous and arboreous plants, belonging to 64 different families (Jolivet, 1988). Most of them are polyphagous, generally feeding on Dicotyledones; several species are reported as pests of wild or cultivated plants. The common reaction to disturbance is the “drop-off reflex”, fol- lowed by a period of thanatosis on the ground. Feedings habits of larvae are very poorly known. They generally live included in a case constructed with faecal matter, hidden on soil surface or under dry leaves. Some of them are myrmecophilous, but most are saprophagous, or free living on leaves. According to our data, there are about 500 described species of Chryptocephalinae in the Palearctic region, belonging to six genera, three of which are reported from Turkey and represented by 91 species, at the present time. | Turkish species have never been studied systematically, although several data and descriptions of new taxa were published in several papers (see Literature cited). In the present work, specimens of most Turkish species have been studied and a list of all known species is given, with taxonomic and biological information. For the first time, an identification key is published for the territory under study. LIST OF THE SPECIES We have examined specimens from the following collections: CMAL: coll. Malmusi, Modena, Italy. CSAL: coll. Saltini, Modena, Italy. HK: coll. Kippenberg, Herzogenaurach, Germany. DE: coll. Erber, Giessen, Germany. ZMH: Zoologiska Museet, Helsingfors Universitet, Finland. JB: coll. Bezdek, Brno, Czech Republic. FK: coll. Kantner, Ceske Budejovice, Czech Republic. MV: Museo civico di Storia naturale, Verona, Italy. MM: Museo civico di Storia natu- rale, Milano, Italy. MZ: coll. Zuber, Kosmonosy, Czech Republic. MNHG: Muséum d'Histoire natu- relle, Geneva, Suisse. SZ: coll. Zoia, Milano. Italy. USR: Dipartimento di Biologia Animale e dell’ Uomo, Università “La Sapienza”, Rome, Italy. Coll. Weise (housed in Museum fiir Naturkunde der Humboldt- Universitàt, Berlin). Coll. Pic (housed in Muséum national d'Histoire naturelle, Paris). Coll. Kraatz (housed in Deutsches Entomologisches Institut, Eberswalde). Coll. Ganglbauer (housed in Naturhistorisches Museum, Wien). Coll. Suffrian (housed in Martin-Luther-Universität, Halle). Lack of abbreviation means that specimens are kept in authors’ collections. The Cryptocephalinae of Turkey TS The species within subgenera are arranged in alphabetical order, as phylogenetic rela- tionships are very poorly known at present. Unfortunately feedings habits and other bio- logical information are unknown for almost all the endemic species. Stylosomus (Stylosomus) flavus Marseul, 1875 Stylosomus Flavus Marseul, 1875: 295. TERRA TYPICA: Greece. GENERAL DISTRIBUTION: Greece; Bulgaria; Caucasus; Turkey. DISTRIBUTION IN TURKEY: Erzurum; Kastamonu; Van (1800m). MATERIAL EXAMINED: Ankara, Beypazari, 10.7.1979, 1 spec.; Artvin, Coruh neheri K., 8.9.1871, 5 specs.; Artvin, Borcka - Artvin, Coruh nehri, 200m, 8.6.86, 1 spec., (MNHG); Kars, Kagizman, Aras nehri, 1200m, 18.6.86, 4 specs., (MNHG); Kars, 20 Km E Kagizman, 1150m, 18.6.86, 3 specs., (MNHG). BIOLOGICAL NOTES: Beetles feed on Tamarix spp. SOURCES: Tomov, 1984; Warchalowski, 1991; Aslan & Ozbek, 1998. Stylosomus (Stylosomus) subelongatus Pic, 1913 Stylosomus subelongatus Pic, 1913: 179. Locus Typicus: Damascus (Syria). GENERAL DISTRIBUTION: Syria; Israel; Turkey. | DISTRIBUTION IN TURKEY: Giimiishane. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: It has been collected on Tamarix spp. SOURCES: Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997a, 1997b. Stylosomus (Stylosomus) tamaricis (Herrich-Schaeffer, 1838) Cryptocephalus tamaricis Herrich-Schaeffer, 1838: 143. TERRA TYPICA: not reported. GENERAL DISTRIBUTION: Southern Europe; Ukraine; Caucasus; Syria; Israel; Central Asia; Northern Africa; Turkey. DISTRIBUTION IN TURKEY: Erzurum; Denizli. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: This species is collected on Tamarix and Myricaria. SOURCES: Sahlberg, 1913; Ogul, 1970; Medvedev & Roginskaia, 1988; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Pachybrachis (Pachybrachis) adaliensis (Weise, 1886) Pachybrachys adaliensis Weise, 1886: 25. Locus TYPICUS: Antalya (= Adalia, TR). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Antalya. MATERIAL EXAMINED: We examined the single spec. in Weise's collection. It is a female, labelled 74 SASSI & KISMALI _as follows: Adalia Korb (handwritten, white label) / 5.1886 (handwritten, white label) / adalien- sis m. (handwritten, white label) / Typus (printed, red label) / Zool. Mus. Berlin (printed, white label) / Pachybrachis adaliensis Wse (handwritten) / L.N. Medvedev det. 1987 (printed). In the original description Weise reported von Heyden as collector of the spec.(s), so the spec. in que- stion would not belong to the type series. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Weise, 1886. Pachybrachis (Pachybrachis) albicans (Weise, 1882) Pachybrachys albicans Weise, 1882: 248 (note). Locus TYPICUS: Kasumkent (near Derbent, Caucasus). GENERAL DISTRIBUTION: Transcaucasus; Turkey. DISTRIBUTION IN TURKEY: Artvin; Erzurum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: It feeds on Salix and Tamarix. SOURCES: Medvedev & Roginskaia, 1988; Warchalowski, 1998. Pachybrachis (Pachybrachis) anatolicus Lopatin, 1985 Pachybrachis anatolicus, Lopatin, 1985: 768. Locus Typicus: Baskale (Van, Turkey). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Van. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. Pachybrachis (Pachybrachis) bodemeyeri (Weise, 1906) Pachybrachys Bodemeyeri Weise, 1906: 472. Locus TYPICUS: Bilecik (Turkey). GENERAL DISTRIBUTION: endemic species from Turkey. DISTRIBUTION IN TURKEY: Bilecik; Erzurum. MATERIAL EXAMINED: “Turchia”, 2 specs., (MV). BIOLOGICAL NOTES: Collected on Quercus spp. (Bodemeyer, 1906) and Salix spp. (Aslan & Ozbek, 1997). SOURCES: Bodemeyer, 1906; Aslan & Ozbek, 1997; Warchalowski, 1998. Pachybrachis (Pachybrachis) excisus (Weise, 1897) Pachybrachys excisus Weise, 1897: 66. Locus TYPICUS: Ankara. GENERAL DISTRIBUTION: Bulgaria; Syria; Turkey. DISTRIBUTION IN TURKEY: Ankara; Antalya. MATERIAL EXAMINED: Kirklareli, Vize, 25.7.1973, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. The Cryptocephalinae of Turkey 75 SOURCES: Gruev & Tomov, 1979; Gruev & Tomov, 1984; Warchalowski, 1991. Pachybrachis (Pachybrachis) fimbriolatus ssp. mendax (Suffrian, 1860) Pachybrachys mendax Suffrian, 1860: 60. TERRA TYPICA: not reported. SYNONYMS: Pachybrachis probus (Weise, 1882); Pachybrachis nitidicollis (Weise, 1894); Pachybrachis dissecticeps (Roubal, 1916). TAXONOMICS REMARKS: Pachybrachis fimbriolatus is a variable taxon with large distri- bution in Palearctic region, and early authors described several related forms whose syste- matic status is not yet clarified up to today. According to the most recent authors, speci- mens from Eastern Europe, Middle Asia and Caucasus might be considered belonging to one (probus: Warchalovski, 1991) or two (probus and nitidicollis: Burlini, 1968) dif- ferent taxa, characterized by broader body, more extended yellow pattern, finer and spar- ser punctures on pronotum, and slight differences on the shape of aedeagus. Lopatin (1977) considered at first the probus and nitidicollis forms as synonyms of the nominal form, but afterwards (1991) he proposed P. probus as a separate species and the ancient . name P. mendax Suffrian as senjor synomym of P. probus. Waiting for further clarifying information, we provisionally assign all Turkish specimens to a single taxon occurring in the oriental part of the range of Pachybrachis fimbriolatus. GENERAL DISTRIBUTION: The real distribution of this subspecies needs further evalua- tion. It has been reported from Moldova; Ukraine; Caucasus; Iran; Central Asia; Southern Russia; Siberia; Mongolia; Turkey. DISTRIBUTION IN TURKEY: Ankara; Bayburt; Bulgar Maden; Cankiri; Denizli; Erzican; Erzurum; Gimiishane; Kars, Sivas; Van (2100m). MATERIAL EXAMINED: Afyon, 1020m, 6.6.1986, 6 specimens; Aksaray, 24.5.61, 1 spec., (MV); Ankara, Beytepe, 30.5.1978, 2 specs.; Ankara, 5-30 Km south of Kirikkale, 1000m, 4.7.1987, 1 spec., (MNUR); Bilecik, Pazaryeri, 25.5.1971, 1 spec.; Bingöl, Kuruca geçidi, 1500-1800m, 24.5.1988, 1 spec.; Bitlis, Karasu cayl., 10 Km W Güroymak, 28.6.1992, 2 specs.; Diyarbakir, Lice, 12.6.1976, 1 spec.; Erzincan, 16.6.1973, 1 spec.; Erzincan, 16.6.1973, 2 spees.; Erzincan, Tercan, 16.6.1973, 1 spec.; Erzincan, Tercan, 1400-1500m, 15.6.1986, 2 specs., (MNUR); Erzincan, Kizildag gecidi, 2150m, 11.7.1975, 1 spec., (MNUR); Erzurum, between Erzurum and Pasinler, 1900m, 7.7.1987, 2 specs., (MNUR); Eskisehir, Hamidiye, 17.5.1979, 2 specs.; Giimiishane, Maden, Kop Dag gecidi, 1700-2300m, 12.7.1987, 3 specs.; Giimiishane, Maden-Bayburt, 1300- 1800m, 16.6.1986, 1 spec.; Istanbul, Beyazit, 12.7.1974, 1 spec.; Konya, Sultan Mts, legit Bodemeyer, 2 specs., (MV); Manisa, Bakir, 11.6.1976, 1 spec.; Sivas, Vassibei gecidi, 1450m, 4.7.1987, 1 spec., (MNUR); Tunceli, 1100m, 15.6.1986, 1 spec., (MNUR). BIOLOGICAL NOTES: Mesophilous and oligophagous species, associated with herbaceous plants belonging to Fabaceae (Onobrychis). Also nominal form has been collected on Fabaceae. Some examined specimens might have been collected on Humulus lupulus and on Salix sp., according to label data. SOURCES: Suffrian, 1860; Escherich, 1897; Bodemeyer, 1900; Sahlberg, 1913; Burlini, 1968; Ogul, 1970; Lopatin, 1977, 1991; Gruev & Tomov, 1979; Tomov, 1984; Aslan & Ozbek, 1997; Aslan & Ozbek, 1998. 76 | SASSI & KISMALI Pachybrachis (Pachybrachis) glycyrrhizae (Olivier, 1808) Cryptocephalus glycyrrhizae Olivier, 1808: 838. Locus TYPICUS: Baghdad (Iraq). GENERAL DISTRIBUTION: Transcaucasus; Syria; Lebanon; Israel; Jordan; Iraq; Iran; Central Asia; Egypt; Afghanistan; Turkey. DISTRIBUTION IN TURKEY: Kars. MATERIAL EXAMINED: Gaziantep, 3.6.1972, 1 spec.; Urfa, Haran, 15.6.1972, 2 specs. BIOLOGICAL NOTES: Probably xerophilous species, associated with the tree layer. It has been collected on Quercus spp., according to Lopatin & Chikanutov (1997); also fee- ding on Glycyrrhiza according to Medvedev & Roginskaia (1998). SOURCES: Berti & Rapilly, 1973; Medvedev & Roginskaia, 1988; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997a, 1997b. Pachybrachis (Pachybrachis) hieroglyphicus (Laicharting, 1781) Cryptocephalus hieroglyphicus Laicharting, 1781: 182. TERRA TYPICA: Tyrol. SYNONYMS: Pachybrachis tristis (Laicharting, 1781); Pachybrachis histrio (Fabricius, 1781); Pachybrachis atomarius (Gebler, 1830); Pachybrachis pseudoscriptus Wagner, 1027 GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus, Central Asia; Western Siberia; Northern Africa; Turkey. DISTRIBUTION IN TURKEY: Bilecik; Konya. MATERIAL EXAMINED: Sivas, Gürün, 19.6.1978, 1 spec.; Sivas, 10.6.1978, 2 specs.; Sivas, 18.6.1970, lispec; BIOLOGICAL NOTES: Hygrophilous species, associated with the tree layer; polyphagous, it feeds on Salicaceae and Betulaceae. SOURCES: Bodemeyer, 1906; Burlini, 1968; Ogul, 1970; Mohr, 1977; Lopatin, 1977; Gruev & Tomov, 1984; Warchalowski, 1991, 1998. Pachybrachis (Pachybrachis) humeralis Burlini, 1956 Pachybrachis humeralis Burlini, 1956: 86. Locus TYPICUS: Bilecik. GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Bilecik. MATERIAL EXAMINED: Karaköse, Tasligay, 6.8.1977, 1 spec.; Kastamonu, 20.6.1974, 1 spec. (uncer- tain determination on a single female). SYSTEMATIC REMARKS: described by Burlini from a couple of specimens labelled by Weise: "var. humeralis" of P. bodemeyeri. It seems to be different from the nominal form, above all by a more extended yellow pattern and shorter and more slender aedea- gus. Further studies on more material are necessary to determine the correct status of this taxon. BIOLOGICAL NOTES: It feeds on Quercus spp., according to Bodemeyer (1906). SOURCES: Bodemeyer, 1906. The Cryptocephalinae of Turkey FF Pachybrachis (Pachybrachis) instabilis (Weise, 1887) Pachybrachys instabilis Weise, 1887: 330. TERRA TYPICA: Kendyk Tau Mts (W. Tian-Shan). GENERAL DISTRIBUTION: Central Asia; Turkey. DISTRIBUTION IN TURKEY: Adapazari; Balikesir; Bursa; Kirklarelı. MATERIAL EXAMINED: none. BIOLOGIC NOTES: This species has been collected on Quercus spp. SOURCES: Medvedev, 1989; Lopatin, 1974; 1977; Aslan & Ozbek, 1997. Pachybrachis (Pachybrachis) laticollis (Suffrian, 1860) Pachybrachyis laticollis Suffrian, 1860: 61. LOCUS TYPICUS: not recorded. SYNONYMS: Pachybrachis misellus (Weise, 1900); Pachybrachis callosus (Sahlberg, 1913). | TAXONOMIC REMARKS: We examined types of P. misellus from Weise’s collection. The series is constituted as follows: 4 specimens (2 males and 2 females) labelled “Mardin” (white label, handwritten), we designated one male as lectotypus of the species; 3 spe- cimens (1 male and 2 females) labelled “Mardin Staund.” (white label, handwritten); 2 specimens (1 male and 1 female) labelled “Zeitun Staund.” (white label, handwritten). GENERAL DISTRIBUTION: Iran; Egypt; Turkey. DISTRIBUTION IN TURKEY: Mardin (as P. misellus). MATERIAL EXAMINED: Corum, Koparan, 25.5.1978, 1 spec.; Diyarbakir, Cermik, 14.6.1972, 1 spec.; Mardin, 7 specs., (coll. Weise); Mardin, 2 specs. (1 “cotypus”), (MV); Sivas, Camlibel, 24.5.1978, 1 spec.; Van, Muradiye, 27.6.1993, 1 spec., (MZ). BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Medvedev, 1989; Lopatin, 1991. Pachybrachis (Pachybrachis) limbatus (Ménétriés, 1836) Cryptocephalus limbatus Ménétriés, 1836: 151. TERRA TYPICA: Greece; Turkey. GENERAL DISTRIBUTION: Balkans; Syria; Israel; Rhodes; Jordan; Turkey. DISTRIBUTION IN TURKEY: Antalya; Balikesir; Bolu; Bursa; Canakkale; Eskisehir (as var. maculatus Suffrian); Golbasi; Istanbul; Izmir; Mersin; Samsun. TAXONOMIC REMARKS: This species is highly variable, perhaps polytypic; at present we are studying the forms attributed to this species. MATERIAL EXAMINED: Antalya, Imrasan Geç. 24.6.97, 2 specs.; Balikesir, Bigadig, 21.4.1975, 1 spec.; Balikesir, Cagis, 380m, 6.6.1986, 1 spec.; Bilecik, Cerkesli, 14.5.1976, 2 specs., (MNHG); Corum, Koparan, 25.5.1978, 2 specs.; Mersin, Namrun, 5.1967 and 5.1997, 3 specs.; Sivas, | spec., (MV). BIOLOGICAL NOTES: Xerophilous species, associated with the tree layer. Probably oli- gophagous (Quercus). Tzanakakis et al. (1963) also reported Pistacia vera and Vitis vinifera as host plants, but in our opinion these data need confirmation. 78 SASSI & KISMALI | SOURCES: Weise, 1889; Bodemeyer, 1900; Sahlberg, 1913; Tzanakakis et al., 1963; Medvedev, 1970; Tomov & Gruev, 1975; Gruev & Tomov, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997. Pachybrachis (Pachybrachis) mardinensis (Weise, 1900) Pachybrachys mardinensis Weise, 1900: 279. Locus TYPICUS: Mardin. SYNONYM: Pachybrachis seidenstueckeri Kippenberg, 1974. GENERAL DISTRIBUTION: Lebanon: caza Bcharrè, Kadicha, 1600m, 1/4.6.1972, 3 speci- mens (new to Lebanon); Syria: Jisr ash Shugur, 26.V.1996, 1 spec. (new to Syria); Israel: Mt. Hermon, 6.V.1986, 1 spec. (new to Israel); Turkey. DISTRIBUTION IN TURKEY: Gaziantep; Mardin. MATERIAL EXAMINED: Adana, Daglari, Hasanbeyli, 27.5.92, 1 spec., (MZ); Adiyaman, Nemrut Mts, 1500-2100m, 13.6.1986, 1 spec., (MNUR); Bitlis, Tatvan, 12.6.93, I spec., (MZ), (uncer- tain identification on 1 female of melanistic form); Eibes (= Hatay, Akbes?), 1 spec., (MV); Elazig, 10 Km N Elazig, 1100m, 14.6.1986, 1 spec., (MNUR); Gaziantep, Nurdagi gec., 1200m, 1.6.92, 1 spec., (MZ); Hakkari, 18 Km East Yiksekova, 30.5.94, 2 specs.; Mersin, Imamli, Silifke, 12.5.94, 1 spec., (MZ), (uncertain identification on 1 female of melanistic form); Mersin, Aydinlar, 1000m, 3 specs. (CMAL); Mus, 1500m, 24.5.1988, 1 spec., (MNUR); Van, Alakabuk Dagi, Kuskuh Kivan, Göründü, 28.6.97, | spec. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Kippenberg, 1974; Lopatin, 1984, 1991; Tomov, 1984; Medvedev, 1989. Pachybrachis (Pachybrachis) nigropunctatus (Suffrian, 1854) Pachybrachys nigropunctatus Suffrian, 1854: 152. GENERAL DISTRIBUTION: Transcaucasus; Iran; Iraq; Syria; Central Asia; Afghanistan. DISTRIBUTION IN TURKEY: Adana. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Feeds on Alhagi and Glycyrrhiza, according to Lopatin (1977). SOURCES: Lopatin, 1977; Aslan & Ozbek, 1998. Pachybrachis (Pachybrachis) pentheri Ganglbauer, 1905 Pachybrachis (Pachystilus) Pentheri Ganglbauer, 1905: 280. GENERAL DISTRIBUTION: Endemic species from Turkey. Locus Typicus: Illany Dagh (Kayseri, Turkey). DISTRIBUTION IN TURKEY: recorded from the type locality only. MATERIAL EXAMINED: 8 syntypes from Ganglbauer collection, labelled as follows: “Illany - Dagh bei Kaisarie” (white label, printed) / “Asia Min. Penther ‘02” (white label, printed) / “Pentheri” (white label, handwritten). We designated a male spec. as lectotype of the species. BIOLOGICAL NOTES: No data were found in the literature. Pachybrachis (Pachybrachis) picus (Weise, 1882) Pachybrachys picus Weise, 1882: 264. TERRA TYPICA: Central Europe. The Cryptocephalinae of Turkey no GENERAL DISTRIBUTION: Central and South Europe; Turkey. DISTRIBUTION IN TURKEY: Erzurum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer; polyphagous it has been collected on Corylus and Rosa spp. SOURCES: Burlini, 1968; Warchalowski, 1991; Aslan & Ozbek, 1998. Pachybrachis (Pachybrachis) scripticollis Faldermann, 1837 Pachybrachis scripticollis Faldermann, 1837: 381. TERRA TYPICA: Armenia. SYNONYM: Pachybrachis israelita Tappes, 1871. GENERAL DISTRIBUTION: Caucasus; Syria; Jordan; Israel; Turkmenistan; Northern Afghanistan; Turkey. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: Diyarbakir, 7.5.1970, 1 spec.; Diyarbakir, Silvan, 27.6.1977, 1 spec.; Diyarbakir, Devegecidi, 17.6.1975, 2 specs.; Gaziantep, Araban, 8.6.1976, 2 specs.; Hakkari, Uludere, 15.6.1976, 13 specs.; Mardin, 3.6.1976, 2 specs.; Suludere, 13.6.1976, 10 specs.; Urfa, Siverek, 7.6.1976 and 16.6.1978, 2 specs.. BIOLOGICAL NOTES: This insect has been collected on Alhagi graecorum according to Lopatin & Chikanutov, 1997. Also sighted on Licorice according with Lopatin (1977). SOURCES: Lopatin, 1977; Lopatin & Chikanutov, 1997. Pachybrachis (Pachybrachis) scriptidorsum Marseul, 1875 Pachybrachis Scriptidorsum Marseul, 1875: 261 (nomen novum). TERRA TYPICA: Caucasus. SYNONYMS: Pachybrachys scripticollis (Suffrian, 1848) nec Faldermann, 1837; Pachybrachis albicans chinensis Weise, 1913. GENERAL DISTRIBUTION: Bulgaria, Bohemia; Ukraine; Caucasus; Syria; Central Asia; Soutwestern Russia; Siberia; Korea; Mongolia; Northern China; Turkey. DISTRIBUTION IN TURKEY: Diyarbakir; Erzurum; Gümüshane. MATERIAL EXAMINED: Artvin, Artvin - Savsat, 500-10.7.1997, 1 spec. BIOLOGICAL NOTES: This insect feeds on Glycyrrhiza, Salix, Populus, Chosenia and perhaps on Myricaria and Statice according to Medvedev & Roginskaia (1988). SOURCES: Burlini, 1968; Ogul, 1970; Tomov & Gruev, 1975; Lopatin, 1977; Gruev & | Tomov, 1984; Medvedev & Roginskaia, 1988; Warchalowski, 1991, 1998; Aslan & Ozbek, 1997a, 1997b. Pachybrachis (Pachybrachis) sinuatus (Mulsant & Rey, 1859) Pachybrachys sinuatus Mulsant & Rey, 1859: 47. Locus TYPpıcus: La Grande-Chartreuse (near Lyon, France). SYNONYM: Pachybrachis haliciensis (Miller, 1868). GENERAL DISTRIBUTION: France, Italy, Central Europe, Balkans; Ukraine; Moldova; 80 SASSI & KISMALI - Caucasus, Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Bolu; Erzurum. The species was also reported from the pro- vince of Antalya: Sogusku-Kizilchamam (Medvedev, 1975) as P. haliziensis [sic], but we could examine one specimen of the mentioned series, which in fact belongs to the closely related species P. velarum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous species, it is probably associated with Salix spp. SOURCES: Burlini, 1968; Medvedev, 1975; Mohr, 1977; Lopatin, 1981; Warchalowski, 1991, 1998; Aslan & Ozbek, 1997. Pachybrachis (Pachybrachis) tessellatus ssp. orientalis (Weise, 1894) Pachybrachys tessellatus var. orientalis Weise, 1894: 144. TERRA TYPICA: Araxes valley. GENERAL DISTRIBUTION: Caucasus; Syria; Israel; Lebanon: Zghorta, Ehden, 1450m, 4.6.1972, 1 specimen (new to Lebanon); Turkey. Nominal form in Southern and Central Europe. DISTRIBUTION IN TURKEY: Adana; Agri; Amasya; Ankara; Bayburt; Bolu; Bursa; Corum; Erzincan; Erzurum; Gaziantep; Giimishane; Kayseri; Mersin; Soguksu; Tokat. MATERIAL EXAMINED: Adana, Erdemli, 15.6.1992, 1 spec., (MZ); Ankara, Akyurt, 15.4.1978, 2 specs.; Antalya, Imrasan, 1300m, 22.6.97, 1 spec.; Antalya, Tepbegasi, 1100m, 11.6.1986, 1 spec., (USR); Corum, Sungurlu-Delece, 800m, 21.6.1986, 1 spec., (USR); Elazig, 10 Km North, 1100m, 14.6.1986, 1 spec., (USR); Erzincan, 5 Km W Alakilise, 1700m, 5.6.1987, 14 specs., (USR); Erzurum, between Erzurum and Pasinler, 1900m, 7.7.1987, 28 specs., (USR); Giimiishane, Maden- Bayburt, 1300-1800m, 16.6.1986, 4 specs., (USR); Giimiishane, 10-30 Km NW Bayburt, 1400- 1800m, 12.6.1987, 6 specs., (USR); Izmir, Agamannon, 10.5.1975, 1 spec., (MNHG); Kahramanmaras, Goksun, 1450-1600m, 18.6.1986, 1 spec., (USR); Kars, 20 Km E Kagizman, 1150m, 18.6.1986, 1 spec., (MNHG); Kars, 10 Km SW Sarikamis, 2000-2100m, 8.6.1987, 4 specs., (USR); Kayseri, Pinarbasi, 18.6.1978, 3 specs.; Makri, 1 spec.; Mardin, Derik, 5.6.1976, 1 spec.; Mersin, Camalan, 11.6.1983, 1 spec.; Nevsehir, 20.6.1968, 8 specs.; Nevsehir, Gore, 23.6.93, 3 specs., (MZ); Sivas, Kizildag gegidi, 1950-2100m, 5.6.1987, 1 spec., (USR); Suhut, 15.6.1972, 1 spec.; Tokat, Almus, 1050-1300m, 19/29.6.94, 1 spec.; Van, Gevas, 29.6.93, 1 spec., (MZ). BIOLOGICAL NOTES: Xerophilous species; nominal form feeds on Quercus spp.; not con- firmed Corylus avellana; unlikely Salix caprea. ‘TAXONOMIC REMARKS: 30% of the specimens examined are totally lacking the yellow spots on pygidium (ab. melanopygus G. Miiller). A large part of the remainder have only signs of this pattern. 70% are lacking the little yellow lateral spot on pronotum. Differences from nominal form seem not to be constant, in our opinion. If the present taxon is confirmed as a true geographic form, it probably should assume the name P. tessellatus ssp. tauricus (Suffrian, 1848). SOURCES: Escherich, 1897; Bodemeyer, 1900; Weise, 1884 , 1906; Ganglbauer, 1905; Medvedev, 1970; Tomov & Gruev, 1975; Tomov, 1984; Warchalowski,1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. The Cryptocephalinae of Turkey 81 X VI, a Figs 1-13: aedeagus. 1-3: Cryptocephalus curda Jacobson (Sivas, Turkey), (scale: 0.62 mm); 4-5: C. lederi Weise (Kabul, Afghanistan); 6-7: C. ocellatus Drapiez (Sondrio, Italy); 8-9: C. tschimganen- sis Weise (from Lopatin, 1977); 10-11: C. pusillus Fabricius (Como, Italy); 12-13: C. tshorumae Tomov (Bolu, Turkey). Figures 4-13: scale: 0.49 mm. 82 SASSI & KISMALI Pachybrachis (Pachybrachis) velarum Warchalowski, 1998 Pachybrachis velarum Warchalowski, 1998: 85. Locus TYPICUS: Refahiye (Erzincan, Turkey). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Erzincan; Erzurum; Gümüshane; Kösk Köyü. MATERIAL EXAMINED: Ankara, Kizilcahamam, 2 spec.; Gümüshane, Maden, Kop Dag gecidi, 1700-2300m, 12.7.1987, 2 specs., (USR); Kars, 16 Km SW Göle, 1600m, 16.6.1986, 26 specs., (MNHG); Kars, 10 Km S Sarikamis, 2000-2100m, 8.7.87, 1 spec., (USR); Kars, Sarikamis, 1.7.1997, 2 specs.; Sivas, Kizildag gecidi, 1950-2000m, 5.7.87, 4 specs., (USR). | BIOLOGICAL NOTES: This species feeds on Salix in mountain and highland regions. Cryptocephalus (Protophysus) moehringi Weise, 1884 Cryptocephalus (Proctophysus) Möhringi, Weise 1884a: 158. Locus Typicus: Amasya (Turkey). SYNONYM: Cryptocephalus bodemeyeri Weise, 1900 n. syn. TAXONOMIC REMARKS: Among 37 specimens received as C. moehringi from the Museum für Naturkunde in Berlin we found the true types of this species mixed with the types of C. bodemeyeri. Some specimens labelled as paralectotypes of C. moehringi are instead syntypes of the latter species. According to the data in the original description (Weise, 1884, 1900), C. moehringi was established on 3 males and 4 females collected by Funcke in Amasya and C. bodemeyri on specimens collected by v. Bodemeyer in Bilecik. Among the material there are 7 specimens (belonging to 3 distinct series, according to label’s data) which were really collected in Amasya. We have kept the foregoing suggestion of Lopatin (unpublished) and designated as lectotype of C. moehringi a male specimen labelled: “Môhringi Amasia” (white label, handwritten) / Lectotypus design. Lopatin 1992 (red label, printed). Another male specimen, with the same data is designated as paralectotype. Three specimens (2 males and 1 female) labelled “Amas” and also label- led as paralectotypes by Lopatin must be rejected because their number does not cor- respond to the original description. Further, 2 specimens (1 male and 1 female) label- led “Amasia Korb” cannot be included as well because the collector does not accord with what the author reported. It follows that 2 specimens constitute the type series of the species. We designated the type series of C. bodemeyeri as follows. Lectotype: 1 female label- led: “Asia Minor Biledjek v. Bodemeyer (printed) / “Proctophysus Bodemeyeri m.” (White label, handwritten). Paralectotypes: 1 male: Asia Minor Biledjek v. Bodemeyer (printed) / “Mit den Typus vergliden Mohr, 1968” / Cryptoc. moehringi Wse det. Mohr / C. Schaefferi möhringi det Lopatin. 4 males and 3 females labelled: “Biledjek v. Bodem.” (white label, handwritten); one of them brings also “Möhringi Ws.” (handw- ritten) and 5 of them had been formerly labelled as paralectotypes of C. moehringi by Lopatin. Anyway, specimens from Amasya and Bilecik are identical in our opinion, so we pro- pose the following new synonymy: Cryptocephalus moehringi Weise, 1884 = Cryptocephalus bodemeyeri Weise, 1900. The Cryptocephalinae of Turkey 83 GENERAL DISTRIBUTION: Albania (?); Greece; Cyprus; Syria; Jordan; Turkey. DISTRIBUTION IN TURKEY: (Some data have been reported as C. bodemeyeri). Amasya; Bilecik; Bolu; Burna; Erzurum; Kars; Mersin. MATERIAL EXAMINED: Ankara, Lalahan, 7.6.1978, 2 specs.; Bilecik, Inegöl-Bozöyük, 750-1000m, 12.6.1991, 1 spec.; Bilecik, Pazaryeri, 7.5.1973, 2 specs.; Bolu, Abant, 1100-1400m, 21.6.1997, 1 spec.; Bolu, Gerede, 28.5.1980, 1 spec.; Bursa, Mudanya, 19.5.1973, 1 spec.; Cankiri, Cerkes, 22.6-4.7.1997 (CMAL); Corum, Alaca, 12.6.1978, 1 spec.; Eskisehir, Porsuk Baraji, 22.5.1998, 4 specs.. BIOLOGICAL NOTES: According to label data of one specimen, it was collected on Crataegus Sp. SOURCES: Bodemeyer, 1900; Weise, 1898; 1900, 1906b; Apfelbeck, 1901; Burlini, 1956; Medvedev, 1970; Tomov & Gruev, 1975; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Protophysus) schaefferi Schrank, 1789 Cryptocephalus Schaefferi Schrank, 1789: 69. TERRA TYPICA: not recorded. SYNONYMS: Cryptocephalus haemorrhoidalis Olivier, 1791; Cryptocephalus unicolor Olivier, 1791; Cryptocephalus notatus Schneider, 1792; Cryptocephalus lobatus Fabricius, 1792. GENERAL DISTRIBUTION: Europe; Balkans; Ukraine; Caucasus; Western Russia; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Erzurum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous speces, associated with the tree layer; polyphagous, it has been collected on Rosaceae, Fagaceae, Corylaceae, Salicaceae. SOURCES: Lopatin, 1977; Aslan & Ozbek, 1997; Warchalowski, 1991; Burlini, 1956. TAXONOMIC REMARKS: It has been reported from Turkey by some authors. We have observed some specimens intermediate with C. moehringi Weise, collected in Southern Balkans, therefore it is possible that C. schaefferi is conspecific with C. moehringi. The latter could represent a geographic variant occurring in the Southeastern part of the range. Further data are necessary in order to solve this question. Cryptocephalus (Protophysus) wehnkei Weise, 1882 Cryptocephalus wehnkei Weise, 1882: 179 (note). TERRA TYPICA: Taurus Mts. GENERAL DISTRIBUTION: It is an endemic species from Turkey. DISTRIBUTION IN TURKEY: Taurus Mts. MATERIAL EXAMINED: Antalya, Seklik, 5.1994, 2 specs., (JB); Mersin, Erdemli, 600-950m, 20.6.93, 2 specs.; Mersin, Namrun, 5.1967 and 5.1997, 2 specs.; Mersin, Namrun, Camliyayla, 2.6.1981, 1 spec. (USR); Imrasan Gec., 24.6.97, 4 specs.; “Taurus”, (Typus ?, Zoologische Museum, Berlin, ex coll. Weise). BIOLOGICAL NOTES: No data were found in the literature. 84 SASSI & KISMALI "TAXONOMIC REMARKS: The single specimen of C. wehnkei in Weise’s collection is a male labelled “Taurus” and “Wehnkei’” and carries two more white labels each with a single unintelligible word that certainly do not correspond to the data of the original descrip- tion. So we consider the specimen as uncertain type. Besides, we had the chance to study female of this species (unknown until now). Dorsal pattern is metallic blue, with yellow spots on apex of elytra, as usual in species of the subgenus Protophysus, but the yellow pattern is more extended anterad, reaching the anterior middle of elytra on sides. Similar widened yellow spots are present in a male specimen as well. Legs of female are metallic blue, but tibiae and femora partly yel- lowish. SOURCES: Weise, 1898. Cryptocephalus (Asionus) amasiensis Weise, 1894 Cryptocephalus amasiensis Weise, 1894: 91. Locus TYPICUS: Amasya. GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: recorded from the type locality only. MATERIAL EXAMINED: Amasya, 23.5.1892, 1 spec., (MV). BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Weise, 1894b; Lopatin, 1965. Cryptocephalus (Asionus) apicalis Gebler, 1830 Cryptocephalus apicalis Gebler, 1830: 201. TERRA TYPICA: Alta] Mts. SYNONYM: Cryptocephalus flavoguttatus Suffrian, 1847. GENERAL DISTRIBUTION: Central and eastern Europe; Balkans; Ukraine; Caucasus; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Turkey, without further informations. MATERIAL EXAMINED: Ankara, Karagöl, Cubuk, 30.6.1979, 1 spec. BIOLOGICAL NOTES: Moderately xerophilous species, associated with herbaceous plants (Artemisia, Potentilla; Ranunculus; Achillea). SOURCES: Clavareau, 1913; Burlini, 1956; Ogul, 1970; Lopatin, 1977; Gruev & Tomov, 1984; Medvedev & Roginskaia, 1988; Warchalowski, 1991. Cryptocephalus (Asionus) curda Jacobson 1897 Cryptocephalus curda Jacobson, 1897: 215. Locus TYPICUS: Kasikoporan (Armenia). SYNONYM: Cryptocephalus araxidis Weise, 1898. GENERAL DISTRIBUTION: Araxes valley; Armenia; Turkey. We have examined several specimens from Khoszov reserve (Armenia), 10.7.1997. DISTRIBUTION IN TURKEY: Afyon; Erzurum; Konya. MATERIAL EXAMINED: Agri, Hamur, 10.8.1974, 2 specs.; Agri, Tendiirek, 2600m, 3.8.1977, 1 spec.; The Cryptocephalinae of Turkey 85 Corum, Iskilip, 6.8.1979, 2 specs.; Erzurum, Horasan, 15.7.1982, 1 spec.; Kars, Igdir, 6.8.1977, 16 specs.; Konya, Aksehir, 1900 (Zoologische Museum, Berlin, ex coll. Weise); Ordu, Kumru, 13.7.1978, 3 specs.; Sivas, Hafik Mt., 1150m and Pasbahce Mt., 1250m, 14.7.79, 6 specs.; betw. Tunceli and Elazig, Yilandag, 11.8.1979, 2 specs.. TAXONOMIC REMARKS: We examined and designated the lectotype of C. araxidis from Weise's collection (Zoologische Museum, Berlin). It is a male specimen, labelled as fol- lows: Caucasus Araxesthal Leder Reitter (printed, white label) / araxidis m. (handw- ritten, white label). We confirm the synonymy Cryptocephalus curda Jacobson, 1897 = Cryptocephalus araxidis Weise, 1898. BIOLOGICAL NOTES: on Verbascum sp., according to label data. SOURCES: Jacobson, 1897; Weise, 1898, 1901; Burlini, 1965; Aslan & Ozbek, 1997a, 1997b. Cryptocephalus (Asionus) pseudoreitteri Tomov, 1976 Cryptocephalus pseudoreitteri Tomov, 1976: 83. Locus TyPICUS: Beypazari (Ankara, Turkey). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: recorded from type locality only. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Tomov, 1976. Cryptocephalus (Asionus) quatuordecimmaculatus Schneider, 1792 Cryptocephalus 14maculatus Schneider, 1792: 195. TERRA TYPICA: Austria. SYNONYMS: Cryptocephalus coloratus Fabricius, 1798; Cryptocephalus pilleri Schrank, 1798. GENERAL DISTRIBUTION: Central Europe; Balkans; Ukraine; Turkey. DISTRIBUTION IN TURKEY: Ankara, Erzurum. | MATERIAL EXAMINED: Ankara, Sögütözü, 1.5.1979, | spec. BIOLOGICAL NOTES: Xerophilous species, feeding on Artemisia, Seseli, Lithospermum, Anthericum. SOURCES: Marseul, 1875; Escherich, 1897; Burlini, 1956; Medvedev & Roginskaia, 1988; Warchalowski, 1991; Aslan & Ozbek, 1998. Cryptocephalus (Lamellosus) angorensis Pic, 1908 Cryptocephalus angorensis Pic, 1908: 14. Locus TYPICUS: Ankara. GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Amasya; Ankara; Corum. Specimens from Amasya and Corum were previously identified as C. laevicollis (Tomov & Gruev, 1975). According to Tomov (1979) they must be attributed to the present species. The old reports of C. laevicollis for Turkey (Weise, 1893) might be referred as well to C. angorensis. 86 SASSI & KISMALI _ MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Pic, 1908; Tomov, 1979. Cryptocephalus (Homalopus) loebli Sassi, 1997 Cryptocephalus (Homalopus) loebli Sassi, 1997: 57. Locus TyPicus: Bolu (Abant, Turkey). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Recorded only for the type locality. MATERIAL EXAMINED: Bolu, Abant, 1500-1600 m; 22.5.76, 2 specs. (typus and paratypus, MNHG); Bolu, Abant, 1.6.1991, 1 spec. (FK); Zonguldak, Safranbolu, 1000m, 5.6.1996, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. Cryptocephalus (Homalopus) prusias Suffrian, 1853 Cryptocephalus Prusias Suffrian, 1853: 95. Locus TYPICUS: Bursa (Turkey). SYNONYM: Cryptocephalus delagrangei Pic, 1898. GENERAL DISTRIBUTION: Syria; Jordan; Turkey. DISTRIBUTION IN TURKEY: Amasya (1000m); Balikesir; Bilecik; Bursa; Erzurum; Giimiishane (1800m); Kirklareli; Mersin; Mus. MATERIAL EXAMINED: Adana, Daglari, Hasanbeyli, 29.5.92, 1 spec., (MZ); Ankara, Kizilcahamam, 9.6.1978, 1 spec.; Bolu, Abant, 28.5.1980, 1 spec.; Cankiri, Cerkes, 22.6-4.7-1997, 1 spec.; Erzurum, Ispir-Camlikaya, 26.6.97, 1 spec. (CSAL); Eskisehir, 4.1934, 2 specs.; Eskisehir, Porsuk Bajari, 22.5.1998, 2 specs.; Konya, Beynam, Giilde, 29.6.1977, 1 spec.; Kiitahya, Gediz, 19.6.1972, 1 spec.; Mersin, Namrun, 5.97, 4 specs.; Sivas, Koyulhisar, 10.6.1991, 1 spec., (JB). BIOLOGICAL NOTES: On Quercus sp. according to Tomov & Gruev (1975). SOURCES: Weise, 1884 ; Pic, 1898; Bodemeyer, 1900; Medvedev, 1970; Tomov & Gruev, 1975; Tomov, 1978; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) anticus Suffrian, 1848 Cryptocephalus anticus Suffrian, 1848: 37. TERRA TYPICA: Caucasus. SYNONYM: Cryptocephalus nigerrimus Ballion, 1878. GENERAL DISTRIBUTION: Caucasus; Syria; Jordan; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Turkey (without further information). MATERIAL EXAMINED: Hakkäri, Yüksekova, 18 Km E, 30.5.94, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Lopatin & Chikanutov, 1997. Cryptocephalus (Cryptocephalus) aureolus Suffrian, 1847 Cryptocephalus aureolus Suffrian, 1847: 132. TERRA TYPICA: Europe. REMARKS: The status of the geographic forms attributed to this species is at present The Cryptocephalinae of Turkey 87 Last | | | 2 N = 23 Figs 14-23: aedeagus. Figs 14-16: Cryptocephalus peyroni Marseul (Hatay, Turkey), (scale: 0.55 mm); 17: C. bicolor Eschscholtz (Astrabad, Iran), (scale: 0.76 mm); 18: C. pelleti Marseul (Lebanon), (scale: 0.76 mm); 19: C. laevicollis Gebler (Moravia), (scale: 0.76 mm); 20: C. angorensis Pic (loca- lity unknown), (scale: 0.59 mm); 21-23: C. volkovitschi Lopatin (Armenia), (scale: 0.76 mm). Figures 17-20 show aedeagus in ventral view. 88 i SASSI & KISMALI under revision by one of us (DS). In this work we prefer to consider C. aureolus as a single widespread and variable taxon. GENERAL DISTRIBUTION: Europe; Caucasus; Jordan; Turkey; Syria; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Antalya; Artvin; Bilecik; Bolu (1400-1600m); Erzincan; Erzurum; Kastamonu (1800m); Kayseri; Manisa (1500m); Probably some of these records might have to be attributed to C. paphlagonius n. sp. MATERIAL EXAMINED: Bolu, Lake Abant, 1500m, 7.1972 and 7.1973, 2 specs.; Erzurum, Kackar Mts, 2000-2500m, 21.7.1983, 1 spec., (HK); Kastamonu, Ilgaz-dag 1800-2200m, 23.7.1963, 1 spec., (DE); Kastamonu, Ilgaz dag, 1800-2300m, 7.1972 and 7.1973, 3 specs.; Kirklareli, Yildiz Mts, Demirköy, 500-900m, 25.6.1997, 2 specs.; Mersin, Camliyayla, 500-800m, 1.6.92, 1 spec.; Yaylak, Kacklar Mt., 2000-2400m, 6/10.7.1997, 5 specs. (JB). BIOLOGICAL NOTES: Polyphagous and meadow mesophilous species, prevalently asso- ciated with Asteraceae Liguliflorae and Ranunculus spp. In Southern part of its range it occurs in mountain and highland regions. SOURCES: Ganglbauer, 1905; Burlini, 1956; Tomov, 1984; Lopatin, 1977; Lopatin & Chikanutov, 1997; Warchalowski, 1991; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) bicolor Eschscholtz, 1818 Cryptocephalus bicolor Eschscholtz, 1818: 466. TERRA TYPICA: not recorded. SYNONYM: Cryptocephalus concinnus Suffrian, 1853. GENERAL DISTRIBUTION: Southern Europe; Ukraine; Balkans; Turkey; Caucasus; Iran; Turkey. DISTRIBUTION IN TURKEY: Erzurum; Tokat. MATERIAL EXAMINED: Kars, 16 Km SW Göle, 1600m, 16.6.1986, 2 specs.. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer (Quercus, Salix and Populus). SOURCES: Ogul, 1970; Rapilly, 1979; Gruev & Tomov, 1984; Medvedev & Roginskaia, 1988; Warchalowki, 1991; E; Aslan & Ozbek, 1997a, 1997b. Cryptocephalus (Cryptocephalus) biguttatus (Scopoli, 1763) Buprestis Biguttata Scopoli, 1763: 65. TERRA TYPICA: not recorded. GENERAL DISTRIBUTION: Europe; Balkans; Ukraine; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Ankara; Bolu (1450-1500m). MATERIAL EXAMINED: Bolu, Abant, 23.6.1969, 9 specs. (USR). BIOLOGICAL NOTES: Mesophilous or hygrophilous species, living on both trees and her- baceous plants as well. According to literature, it seems to be polyphagous and has been collected on species of several plant families (Salicaceae, Fagaceae, Corylaceae, Betulaceae, Ericaceae, Asteraceae and Juncaceae). SOURCES: Ogul, 1970; Tomov & Gruev, 1975; Lopatin, 1977; Tomov, 1984; Gruev & Tomov, 1984; Warchalowski, 1991. The Cryptocephalinae of Turkey 89 Cryptocephalus (Cryptocephalus) biledjekensis Pic, 1909 Cryptocephalus cribratus var. biledjekensis Pic, 1909: 153. Locus TYPICUS: Bilecik. GENERAL DISTRIBUTION: Bulgaria; Turkey. DISTRIBUTION IN TURKEY: Bilecik; Gümüshane. MATERIAL EXAMINED: Ankara, Kizilcahamam, 12.7.1979, 1 spec.; Izmir, 1 spec.; Izmir, Bergama, 14.5.1971, 1 spec.; Konya, Dozar, 1 spec.; Siirt, Aktas, 650m, 20.5.1988, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Pic, 1909; Tomov, 1978; Gruev & Tomov, 1984; Warchalowski, 1991. Cryptocephalus (Cryptocephalus) bipunctatus (Linné, 1758) Chrysomela 2-punctata Linné, 1758: 374. TERRA TYPICA: Europe. SYNONYM: Cryptocephalus dispar Paykull, 1799. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Siberia; NE China; Korea; Turkey. The ancient report for Northern Africa (Weise, 1893) has never been confirmed. DISTRIBUTION IN TURKEY: Amasya; Ankara; Artvin; Bolu; Bursa; Canakkale; Corum; Erzincan; Erzurum; Izmir; Kars; Mersin; Sinop; Sivas (1650m). MATERIAL EXAMINED: Ankara, Kizilcahamam, 6.1992,1 spec., (JB); Bolu, Abant, 1100-1400, 21.6.97, 2 specs. (CSAL); Bursa, 19.6.1967, 2 specs.; Corum, Bogazkale, 1150m, 10.7.1995, 1 spec. (USR); Istambul, Alem Dagh, 10.6.1966, 1 spec.; Izmir, Yeniköy, 60m, 3.6.1991, 1 spec.; Kars, Sarikamis, 2000-2200m, 8.7.1987, 6 specs.; Kars, Sarikamis, 2000m, 21.6.1994 and 7.6.1998, 6 specs.; Kars, Sarikamis, 28.6.97, 2 specs. (CSAL); Kirklareli, Lüleburgaz, 29.5.91, 2 specs., (JB); Konya, 15.6.1980, 3 specs.; Konya, Aksehir, 6.1967, 1 spec.; Kiitahya, Orencik, 20.5.1971, 1 spec.; “Lycia”, 2 specs.; Manisa, Sabuncubeli, 10.6.1972, 1 spec.; Mersin, Camliyayla, | spec., (CMAL); Mersin, Camliyayla, 1400-1850m, 9.6.1986, 1 spec.; Mersin, Camliyayla, 29.6/3.8.1997, 4 specs., (JB); Mersin, Guzeloluk, 25.6.97, 1 spec., (CHIARI): Tokat, Artova, 2.7.1978, 1 spec.; Zonguldak, Caycuma, 16.7.1979, 1 spec. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous (Salicaceae, Fagaceae, Corylaceae and Betulaceae). Some specimens were collected on Paliurus sp. SOURCES: Escherich, 1897; Sahlberg, 1913; Burlini, 1956; Gressitt & Kimoto, 1961; Bodemeyer, 1900; Medvedev, 1970; Ogul, 1970; Tomov & Gruev, 1975; Lopatin, 1977; Tomov, 1984; Aslan & Ozbek, 1997 Aslan & Ozbek, 1998. Cryptocephalus (Cryptocephalus) cordiger (Linné, 1758) Chrysomela cordigera Linné, 1758: 375. TERRA TYPICA: Europe. GENERAL DISTRIBUTION: Central and Eastern Europe; Balkans; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Erzurum. MATERIAL EXAMINED: none. 90 SASSI & KISMALI BIOLOGICAL NOTES: Polyphagous and mesophilous, associated with the tree layer (Fagaceae, Betulaceae, Salicaceae and Rosaceae). SOURCES: Lopatin, 1977; Warchalowski, 1991; Aslan & Ozbek, 1998. Cryptocephalus (Cryptocephalus) crassus Olivier, 1791 Cryptocephalus crassus Olivier, 1791: 620. TERRA TYPICA: Var (France). SYNONYM: Cryptocephalus gravidus Herrich-Schäffer, 1838. GENERAL DISTRIBUTION: Southern Europe; Syria; Israel; Northern Africa; Turkey. Reports from Caucasus and Turkmenistan (Marseul, 1875) have not been confirmed by more recent authors. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Xerophilous species, moderately polyphagous, associated with shrubby Papilionaceae (“Brooms”) and Asteraceae (Artemisia). Feeding on Artemisia monosperma in eastern part of its range according with Lopatin & Chikanutov (1997). SOURCES: Marseul, 1875; Lopatin & Chikanutov, 1997. Cryptocephalus (Cryptocephalus) cribratus Suffrian, 1847 Cryptocephalus cribratus, Suffrian, 1847: 90. Locus TYPICUS: Istanbul. SYNONYM: Cryptocephalus trapezensis Tappes, 1871 n. syn. TAXONOMICS REMARKS: Thanks to the kindness of Dr Nicole Berti, we were able to exa- mine the single specimen of C. trapezensis Tappes in Tappes’ collection. It is a fema- le, labelled as follows: “Persath Deyr” (printed, white label) / “597” (handwritten, white label) / “spec ?” (handwritten, white label) / “42” (handwritten, white label) / “type” (printed, red label)/ “Cryptocephalus trapezensis Tappes” (handwritten, white label ). Data collection do not completely agree with what the author reported in the descrip- tion: indeed, “Persath” is quite an enigmatic information and it seems not to correspond with any of the present Turkish toponyms (type locality is Trabzon). On the other hand the collector (Deyrolle) is the same. Anyway we think that the specimen properly repre- sents the sense that Tappes gave to the species, so we designated it as neotype of the species. In our opinion Cryptocephalus trapezensis should be considered a chromatic form of C. cribratus. The latter is a variable species and all intermediates have been observed. Furthermore, the aedeagus of the trapezensis form does not differ signifi- cantly. Accordingly, we propose the following synonymy: Cryptocephalus cribratus Suffrian, 1847 = Cryptocephalus trapezensis Tappes, 1871. GENERAL DISTRIBUTION: Caucasus; Iran; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Bilecik; Erzurum (as C. trapezensis); Gümüshane (1800m); Konya; Trabzon; (as C. trapezensis). MATERIAL EXAMINED: “Anatolia, ab. anatolicus mihi”, 1 spec., (MV) (as C. trapezensis); Ankara, 14.5.1980, 1 spec.; Ankara, Karagöl, Cubuk, 20.6.1979, 1 spec.; Bilecik, leg. Bodemeyer, 2 specs., (MV); Bolu, Abant, 1100-1400m, 21.6/5.7.1997, I spec.; Erzurum, Ilica, Yildizeli, Sivas, 1250m, The Cryptocephalinae of Turkey CH 14.7.1979, 1 spec.; Erzurum, Kackar Mt., Yaylalar, 5/10.7.1997, 5 specs., (JB); Izmir, Bademli, 5.6.1969, 1 spec.; Kars, Sarikamis, 2000m, 7.6.1998, 1 spec.; Nevsehir, Urgiip, 20.5.1978, 1 spec. BIOLOGICAL NOTES: On Populus nigra, according to label data. Feeding on Salix sp., after Aslan & al. (1997). SOURCES: Tappes, 1871; Bodemeyer, 1900; Weise, 1901; Tomov & Gruev, 1975; Lopatin, 1977; Tomov, 1979; Rapilly, 1980; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) duplicatus Suffrian, 1847 Cryptocephalus duplicatus Suffrian, 1847: 126. TERRA TYPICA: Caucasus. SYNONYM: Cryptocephalus unicolor Faldermann, 1837 nec Olivier, 1791. GENERAL DISTRIBUTION: SE Bulgaria; Caucasus; Jordan, Israel; Turkey. DISTRIBUTION IN TURKEY: Several authors reported the present species under the name C. concolor Suffrian, 1847. According to Lopatin (1984) C. concolor does not reach the territory under study. Adapazari; Amasya; Ankara (1300m); Antalya (2000m); Artvin (2200m); Bilecik; Bitlis (2300m); Bolu; Bursa (1400m); Corum (1150m); Erzincan (2150m); Erzurum (2300m); Giresun (1800-2000m); Gümüshane; Isparta; Izmir; Izmit; Kars (2000m); Kastamonu (1800m); Konya (1200m); Mersin; Samsun; Sinop; Sivas (1650m); Tunceli (900m); Trabzon; Van; Yozgat (1300m). MATERIAL EXAMINED: Adana, Osmaniye, Hasanbeyli, 5.1970 1 spec., (HK); Adapazari, Sapanca, 17.5.1973, 1 spec.; Amasya, Aydinca, 600m, 24.6.1997, 2 specs. (CSAL); Amasya, Inegöl Dagi, 900m, 23.6.1997, 1 spec. (CSAL); Ankara, 9.6.1977, 1 spec.; Ankara, Ayas, 10.7.1979, 3 specs.; Ankara, Beynam, 26.6.1977, 2 specs.; Ankara, Cogun, 23.6.1993, 1 spec., (MZ); Ankara, Karagöl, 7.7.1978 1 spec.; Ankara, Kizilcahamam, 25.5-6.7.1970, 1 spec., (HK); Ankara, Cogun Lake, 22.6.1993, 2 specs., (MZ); Antalya, Giindogmus, 27.6.1980 and 23.6.1997, 2 specs.; Antalya, Imrasan Gec., 24.6.1997, 2 spec.; Antalya, Sögüt, 20.6.1981, 1 spec.; Antalya, Yarpuz, 12.6.1997, 5 specs.; Artvin, 12.6.1973, 4 specs.; Artvin, Ardanuc, 1550m, 10.6.1998, 2 specs.; Artvin, Artiparmak, Yusufeli, 1400-1800m., 20.7.1983, 2 specs., (DE); Artvin, Artvin - Savsat, 500/1000m, 26.6.1997, 3 specs., (CSAL); Artvin, Bilbilan, Yalnizcam Pass., 1700-2600m, 16 specs., (DE); Artvin, Kackar Mt., Yaylak, 2000-2400m, 6/10.7.1997, 9 specs. (JB); Artvin, Kackar Mt., Yaylalar, 5/10.7.1997, 23 specs., (JB); Artvin, Kafkasoryayla, 1600-1700m, 5.7.1992, 1 spec., (DE); Artvin, Karafsor, 23.6.1990, 3 specs., and 6.7.1993, 2 specs., (DE); Artvin, Karçkal dagh, Otingöl, 2000- 2600m, 10.7.1983, 2 specs., (DE); Artvin, Savsat-Carn Geg., 1000/1500m, 26.6.1997, 1 spec. (CSAL); Artvin, Sarigöl, Biçakçilar, 13.7.96, 1 spec., (MZ); Bilecik, 17.7.1972 and 15.6.1977, 3 specs.; Bilecik, Gölpazari, Besevler, 450 m., 23.5.1996, 1 spec., (MZ); Bilecik, Pazaryeri, 16.7.1972, 1 spec.; Bilecik, Sögüt, 20.6.1975 and 20.6.1981, 3 specs.; Bolu, Abant, 1100-1400m, 21.6.1997, 1 spec., (CMAL); Bolu, Seben, 12.7.1978, 4 specs.; Bursa, Karacabey, 15.6.1976, 2 specs.; Bursa, Cagliyan, 19.7.1997, 10 specs., (JB); Bursa, Uludag, Bursa-Sugukpinar, 1000m., 19.7.1989, 1 spec., (DE); Bursa, Uludag, 1800m, 24.7.1988, 1 spec., (HK); Cankiri, Cerkes, 4.7.1997, 4 specs. (CSAL); Cankiri, Cerkes, 22.6.1997, 2 specs.; Cankiri, Eskipazar, 31.7. 1979, 1 spec.; Corum, 6.1973 and 6.1974, 2 specs.; Corum, Kos Mt., 1100-1900m., 1 spec., (DE); Eregli, 25.5.1980, 1 spec.; Erzincan, Tercan, 1400m, 20.6.1994, 1 spec.; Erzurum, Dumlu-Koyu, 2000m., 28.7.1973, 1 spec., (DE); Erzurum, Kackar Mt., Hevek, 2000-2500m, 7.8.1984, 2 specs.; Erzurum, Kackar Mt., Hevek, (Meretet-yayla), 2000-2500m 1 spec., (HK); Erzurum, Pazaryolu-Golyurt Gec., 1500-2000m, 4 specs.; Eskisehir, Mahmudiye, 8.7.1979, 2 specs.; Gindrak, 4.6.1971, | spec.; Giimiishane, 11.7.1978, 2 specs.; Giimiishane, Kilickaya, 1700-2100m, 17.7.1992, 3 specs., 92 SASSI & KISMALI (DE); Gümüshane, Maden, Kop Dag gecidi, 2200-2500m, 5.8.1995, 14 specs. and 29.7.1993 1 spec., (DE); Gümüshane, Maden, Kop Dag gecidi, 2200-2500m, 19.7.1976 1 spec., (HK); Hakkari, Cukurca, 14.6.1976, 2 specs.; Isparta, Egridir, Kovada, 5.73, 1 spec., (HK); Kahramanmaras, Maras, 11.6.1985, 1 spec.; Karaman, 2.7.1980, 1 spec.; Kars, Ardahan, 6.7.1975, 4 specs., (DE); Kars, Ardahan, Posof, 29.6.1993, 2000m, 1 spec., (DE); Kars, Ardahan, Hanak, 1900-2100m, 7.8.1970, 2 specs., (DE); Kars, Karakurt, 1.7.97, 3 specs. (CSAL); Kars, Sarikamis, 28.6.1997, 7 specs. (CSAL); Kars, Sarikamis, 2000m, 21.6.1994 and 7/11.6.1998 8 specs.; Kars, Sarikamis, 2000-2200m, 26.7.1983 and 3.8.1985, 4 specs., (DE); Kars, Sarikamis, 15.7.1982, 2 specs.; Kastamonu, Cide, 9.6.1980, 2 specs.; Kastamonu, Ilgaz Mts, 7.8.1996, 2 specs., (JB); Kirklareli, Lüleburgaz, 29.5.1991, 4 specs., (JB); Kirklareli, Yildiz Mts, Demirköy, 500-900m, 25.6.1997, 2 specs.; Artvin, Kobak, Sarigöl, 14.7.1996, 3 specs., (MZ); Kojedagi gec., 28.6.1994, 14 specs.; Konya, Beysehir Lake, 1.8.1996, 1 spec., (JB); Konya, Doganhisar, 5.7.1980, 2 specs.; Konya, Hadim, 4.7.1980, 1 spec.; Konya, Koghisar, 29.5.1973, 1 spec.; Konya, Sultan Mts, Cankurtaran, 1800m, 26.5.1996, 1 spec., (MZ); Konya, Sultan Mts, Cay, 1400-2000m., 15.8.1971, 3 specs.; Köseköy, 22.5.1995, 2 specs., (JB); Kütahya, 19.6.1975, 1 spec.; Manisa, Derbent, Dramon, 6.6.1991, 4 specs., (MZ); Mersin, Aydinlar, 1000m, 1 spec. (CMAL); Mersin, Camliyayla, 29.6.1997, 5 specs., (JB); Mersin, Camliyayla, Sebil, 2.6.1995, 1 spec., (MZ); Mersin, Camliyayla 1000-1400m, 3 specs. (CMAL); Mersin, Erdemli, 600-950m, 20.6.1993, 13 specs.; Mersin, Erdemli, Kayaci, 2.5.1994, 1 spec., (JB); Mersin, Namrun, 5.1967, 1 specs.; Mersin, Tarsus, Gamlagaye, 1 spec. (MM); Mus, Buglan gec., 1650m, 2.6.1998, 1 spec.; Nevsehir; Hacibektas, 26.6.1980, 1 spec.; Ordu, Persembe, 8.6.1973, 1 spec.; Samsun, Bafra, 4.6.1973, 5 specs.; Samsun, Karadag geç., 700-900m, 26.6.1983, 1 spec., (HK); Samsun, Karadag Geg., 26.6.1983, 2 specs.; Samsun, Karadag Geg., 700-900m, 26.4.1983, 6 specs., (DE); Samsun, Kavac-Hacilar Gec., 24.6.1997, 4 specs.; Samsun, Ladik, 1.7.1978, 1 spec.; Sarica, 1500m, 9.8.1985, 1 spec., (DE); Surt, Kurtalan, 23.6.1976, 1 spec.; Sivas, Serefiye, 1900m., 29.6.1992, 4 specs., (DE); Sivas, Zara - Serefiye, 1600m., 23.6.1987, 3 specs., (DE); Tekirdag, 1.6.1978, 8 specs.; Tokat, Almus, 1050-1300m, 29.6.1994, 1 spec.; Tokat, Camlibel Gec., 1600m., 22.6.1987, 1 spec., (DE); Tokat, Zile, 3.7.1978, 1 spec.; Trabzon, 9.6.1973, 1 spec.; Trabzon, Turul, 4.7.1993, 5 specs.; Trabzon, Zigana, 8.7.1978, 2 specs.; Van, Kuzgunkiran Geg., 1.7.1997, 2 specs., (JB); Van, Kuzgunkiran, Gevas, 2300m, 23.7.1985, 1 spec., (DE); Yozgat, Sarikaya, 14.7.1963, 1 spec.; Zonguldak, Caycuma, 16.7.1979, 1 spec.; Zonguldak, Ulus, 17.7.1979, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. The species probably feeds on weeds (Asteraceae?). We can assume that the present species has affinities with C. hypo- chaeridis s. 1. in regards to the ecological features. SOURCES: Weise, 1893; Escherich, 1897; Bodemeyer, 1900; Berti & Rapilly, 1973; Tomov & Gruev, 1975; Medvedev, 1983; Tomov, 1984; Gruev & Tomov, 1984; Lopatin, 1981, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) flavipes Fabricius, 1781 Cryptocephalus flavipes Fabricius, 1781:146. TERRA TYPICA: Italy. SYNONYM: Cryptocephalus wydleri Faldermann, 1837. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Israel; Iran; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Amasya; Artvin; Aydin; Bolu; Bursa; Canakkale; Erzurum; Eskisehir; Gaziantep; Istanbul; Kastamonu; Kars; Malatya; Mersin; Ordu; Samsun; The Cryptocephalinae of Turkey 95 Sinop; Tokat; Tunceli; Trabzon. Material examined: Amasya, Merzifon, 25.5.1978, 2 specs.; Ankara, Karagöl, 7.7.1978, 1 spec.; Artvin, Yalnizcam geg., 2700m, 10.6.1998; Bolu, Mengen, 26.5.1980, 1 spec.; Bursa, Uludag, 31.5.1971, 1 spec.; Cankiri, Cerkes, 22.6.1997, 1 spec.; Izmit, 25.4.1975, 1 spec.; Kars, Sarikamis, 2000m, 7.6.1998,4 specs.; Mersin, Camliyayla, 1300m, 1.7.1993, 2 specs.; Mersin, Namrun, 5.1967, 3 specs.; Rize, Ikizdere, 500-1000m, 25.7.1997, 1 spec.; Rize, Ikizdere, 1000-1500m, 25.6.1997, 1 spec.; Tokat, Almus, 2.7.1978, 1 spec.; Yozgat, Cökerek, 13.6.1978, 1 spec. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous, it is found routinely on Salicaceae, Corylaceae, Fagaceae, Rosaceae and Betulaceae. SOURCES: Bodemeyer, 1900; Medvedev, 1970; Ogul, 1970; Berti & Rapilly, 1973; Tomov & Gruev, 1975; Lopatin, 1977; Tomov, 1984; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) fulmenifer Reitter, 1889 Cryptocephalus fulmenifer Reitter, 1889: 376. TERRA TYPICA: Caucasus. SYNONYMS: Cryptocephalus undatus Suffrian, 1853; Cryptocephalus phaleratus Suffrian, 1853; Cryptocephalus tigrinus Berti & Rapilly, 1979. GENERAL DISTRIBUTION: Caucasus; Iran; Turkey. DISTRIBUTION IN TURKEY: Araxes valley. MATERIAL EXAMINED: Adiyaman, Karadut, Nemrut Mts, 1700m, 9/12.7.1997, 2 specs., (JB); Diyarbakir (Eläzig), Ergani - Maden, 3.9.1939, 1 spec., (MNHG). BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Berti & Rapilly, 1979; Lopatin, 1985. Cryptocephalus (Cryptocephalus) gloriosus Mulsant, 1853 Cryptocephalus gloriosus Mulsant, 1853: 159. TERRA TYPICA: Karaman (Turkey). TAXONOMIC REMARKS: We could not examine types of this species. According to Lopatin (1965) it may belong to the subgenus Asionus. GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: The species has been reported from the type locality only. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Marseul, 1875. Cryptocephalus (Cryptocephalus) ilicis Olivier, 1808 Cryptocephalus ilicis Olivier, 1808: 801. TERRA TYPICA: "Orient". SYNONYMS: Cryptocephalus siculus Herrich-Schäffer, 1836. GENERAL DISTRIBUTION: Jordan; Syria; Balcan Peninsula; Sicily; Turkey. DISTRIBUTION IN TURKEY: Balikesir; Istanbul. 94 SASSI & KISMALI MATERIAL EXAMINED: Bursa, 18.5.1973, 1 spec.; Izmir, Bergama, 1.5.1990, 1 spec., (JB); Izmir, Bayindir, 24.4.1973, 2 specs.; Kirklareli, Liileburgaz, 8.5.1975, 2 specs.; Konya, Sultanyayla, 2.6.1972, 1 spec.; Manisa, Manisa Mt., 5.1994, 1 spec., (JB); Ores, 6.7.1982, 2 Specs.. BIOLOGICAL NOTES: On Crataegus sp., according to label data. Probably associated with the tree layer (Quercus spp.). SOURCES: Pic, 1900, 1908; Ganglbauer, 1905; Sassi, 1995; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) imperialis Laicharting, 1781 Cryptocephalus imperialis Laicharting, 1781; 179. Locus TYPICUS: Bolzano (Italy). SYNONYM: Cryptocephalus bistripunctatus Germar, 1824. GENERAL DISTRIBUTION: Europe; Caucasus; Iran; Turkey. DISTRIBUTION IN TURKEY: Amasya; Erzurum; Konya. MATERIAL EXAMINED: Usak, 12.6.1972, 1 spec. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous, on Corylaceae, Betulaceae, Fagaceae and Rosaceae. SOURCES: Weise, 1901; Medvedev, 1975, 1983; Medvedev & Roginskaia, 1988; Aslan & Ozbek, 1997, Cryptocephalus (Cryptocephalus) janthinus (Germar, 1824) Cryptocephalus janthinus Germar, 1824: 555. TERRA TYPICA: Germany. SYNONYMS: Cryptocephalus megaloderus Faldermann, 1830; Cryptocephalus rusticus Faldermann, 1837. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Iran; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Canakkale; Eskisehir; Mersin. MATERIAL EXAMINED: Eskisehir, Boziiytik, 17.8.1967, 2 specs. (USR); Kumlona, 28.8.1979, 1 spec.; Kirklareli, Pinarhisar, 7.7.1972, 1 spec. BIOLOGICAL NOTES: Meadow hygrophilous species; probably polyphagous. Lysimachia vulgaris, Lythrum salicaria, Phragmites communis are probable host plants. SOURCES: Ogul, 1970; Berti & Rapilly, 1973; Tomov & Gruev, 1975; Tomov, 1984; Lopatin, 1977; Warchalowski, 1991. Cryptocephalus (Cryptocephalus) moraei (Linné, 1758) Chrysomela Moraei Linné, 1758: 374. TERRA TYPICA: Europe. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Iran; Jordan; Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Bilecik; Bolu (1400-1600m); Bursa; Diyarbakir; Erzincan; Erzurum; Giresun; Gümüshame; Izmir; Kars; Kastamonu (1800-2300m); Samsun. MATERIAL EXAMINED: Adana, Bahce, 7.6.1984, 1 spec.; Adana, Gulek, Derekenar, 7.6.1978, 1 The Cryptocephalinae of Turkey 95 Figs 24-26; 28-30; 32-34: aedeagus, (scale: 1.61 mm). Figs 27; 31; 35: abdomen in ventral view, (scale: 2.82 mm). 24-27: Cryptocephalus euchirus Kraatz (Iran); 28-31: C. moehringi Weise (Amasia, Turkey, Lectotypus); 32-35: C. wehnkei Weise (Imrasan gegidi, Antalya, Turkey). 96 SASSI & KISMALI spec.; Adapazari, 15.7.1972, 1 spec.; Adapazari, Hendek, 14.7.1972, 8 specs.; Adapazari, Sapanca, 15.7.1972, 2 specs.; Afyonkarahisar, Sincanli, 13.6.1972, 1 spec.; Akarsen, 1500m, 8.8.1971, 1 spec.; Ankara, Karagöl, Cubuk, 30.6.1979, 1 spec.; Artvin, 12.6.1978, 1 spec.; Bitlis, Tatvan, 9.7.1977, 1 spec.; Bolu, Mengen, 14.8.1979, 1 spec.; Bursa, 30.5.1971, 18.7.1972 and 20.5.1973, 3 specs.; Bursa, Inegöl, 19.6.1975, 2 specs.; Bursa, Orhangazi, 2.6.1971, 1 spec.; Bursa, Uludag, 18.7.1972, 1 spec.; Eregli, 16.7.1979, 1 spec.; Giimiishane, 11.7.1978, 1 spec.; Hakkari, Cukurca yol ayrimi, 11.6.1975, 1 spec.; Izmir, Agamemnon, 10.5.1975, 2 specs.; Karatas, 5.1967, 1 spec.; Kars, Sarikamis, 2000-2100m), 8.6.1987, 1 spec.; Kirklareli, 24.7.1973, 2 specs.; Konya, Doganhisar, 5.7.1980, 2 specs.; Mersin, Camliyayla, 29.6.1997, 2 specs., (JB); Mugla, Seki, 9.6,1973, 2 specs.; Ordu, Kumru, 13.7.1978, 1 spec.; Rize, Hayrat, 10.6.1973, 1 spec.; Samsun, Bafra, 4.6.1973, 3 specs.; Rize, 10.6.1973, 1 spec.; Samsun, Kavac, Hacilar Geç., 24.6-3.7.1998, 1 spec., (CSAL); Trabzon, 12.7.1972, 1 spec.; Trabzon, Macka, 27.6.1997, 1 spec., (JB); Tunceli, 1100m, 15.6.1986, 1 spec.; Zonguldak, Caycuma, 31.5.1980, 1 spec.; Zonguldak, Ulus, 29.5.1980, 1 spec. BIOLOGICAL NOTES: Meadows mesophilous species; oligophagous, it probably feeds only on Hypericum spp. (Guttiferae). SOURCES: Sahlberg, 1913; Burlini, 1956; Ogul, 1970; Tomov & Gruev, 1975; Gruev & Tomov, 1979; Rapilly, 1980; Tomov, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) octacosmus Bedel, 1891 Cryptocephalus octacosmus Bedel, 1891: 131, nomen novum. TERRA TYPICA: Toscana (Italy). SYNONYM: Cryptocephalus sexpustulatus Rossi, 1790: 97, nec Villers, 1789: 156. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Western Siberia; Western China; Turkey. DISTRIBUTION IN TURKEY: Ankara; Bayburt; Bilecik; Bolu; Denizli; Erzurum; Eskisehir; Isparta; Izmir (as C. sexpustulatus Rossi, 1790); Konya; Tokat. MATERIAL EXAMINED: Adana, Tufanbeyli, 5.6.1984, 2 specs.; Adapazari, Sapanca, 15.7.1972, 1 spec.; Antalya, Saklikent, 1860m, 26.7.1985, 1 spec.; Bolu, Seben, 12.7.1978, 5 specs.; Bursa, Cagliyan, 10/14.7.1997, 1 spec., (JB); Cankiri, Cerkes, 23.7.1979, 1 spec.; Diyarbakir, 12.6.1972, 2 specs.; Eskisehir, Mahmudiye, 8.7.1979, 1 spec.; Gümüshane, Maden, Kop Dag gecidi, 1700- 2300m, 12.7.1987, 1 spec.; Isparta, Anamas, 3 specs.; Kayseri, Biinyan, 19.6.1978, 1 spec.; Kirklareli, Liileburgaz, 1.6.1975, 1 spec.; Manisa, Bakir, 12.6.1972 and 11.6.1976, 2 specs.; Mardin, Emerli, 12.6.1972, 2 specs.; Mus, Karakiitiik, 12.6.1975, 2 spec.; Mus, Solhan, 15.6.1975, 1 spec.; Nevsehir, Urgiip, 29.5.1973, 1 spec.; Nigde, Camardi, 29.6.1980, 1 spec.; Nigde, Ulukisla, 1.7.1980, 2 specs.; Pinarhisar, 7.7.1972 and 3.6.1975, 3 specs.; Samsun, Ladik, 1.7.1978, 2 specs.; Sivas, 18.6.1978, 1 spec.; Sivas, Ilica, Yildizeli, 14.7.1979, 2 specs.; Tekirdag, 1.6.1975, 1 spec.; Tunceli, Pulumur gegidi, 1350-1900m, 6.7.1987, 1 spec.; Zonguldak, Bartin, 31.5.1980, 2 specs.. BIOLOGICAL NOTES: The old data for Fraxinus, Alnus and other trees and bushes (Marseul, 1875) should be dismissed. It is a meadow hygrophiious species and Sanguisorba offi- cinalis is presently the only ascertained food-plant (Duhaldeborde, 1994). SOURCES: Escherich, 1897; Weise, 1901; Sahlberg, 1913; Burlini, 1956; Ogul, 1970; Medvedev, 1973; Tomov & Gruev, 1975; Medvedev, 1975; Lopatin, 1977; Tomov, 1984; Warchalowski, 1991; Duhaldeborde, 1994; Aslan & Ozbek, 1997. The Cryptocephalinae of Turkey 97 Cryptocephalus (Cryptocephalus) octomaculatus Rossi, 1790 Cryptocephalus octomaculatus Rossi, 1790: 96. TERRA TYPICA: Toscana (Italy). SYNONYMS: Cryptocephalus quinque punctatus Harrer, 1784 nec Scopoli, 1763; Cryptocephalus 12 punctatus Fabricius, 1792. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Southern Siberia; Turkey. DISTRIBUTION IN TURKEY: Konya. MATERIAL EXAMINED: Konya, Aksehir, 1200m, 5.8.1967, 2 specs., (USR); Mersin, Namrun, 35 Km NNW of Tarsus, 20/22.7.1997, 1 spec., (JB). BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. This species feeds on Quercus spp. Older authors reported several other genera which need confir- mation (Carpinus, Corylus and Populus). SOURCES: Ogul, 1970; Tomov, 1984; Warchalowski, 1991; Mohr, 1997. Cryptocephalus (Cryptocephalus) octopunctatus (Scopoli, 1763) Buprestis Octopunctata Scopoli, 1763: 67. TERRA TYPICA: not recorded. SYNONYMS: Cryptocephalus variabilis Schneider, 1791; Cryptocephalus potaissiae Csiki, 1953. GENERAL DISTRIBUTION: Europe; Ukraine; Central Asia; Western Siberia. DISTRIBUTION IN TURKEY: Ankara. MATERIAL EXAMINED: Adapazari, 1924, (MV); Tokat, Almus 1050-1300m, 19/29.6.94 1 spec. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous, on Salicaceae, Betulaceae, Rosaceae and Fagaceae. SOURCES: Escherich, 1897; Ogul, 1970; Lopatin, 1977; Gruev & Tomov, 1984. Cryptocephalus (Cryptocephalus) paphlagonius n. sp. The species is very similar to Cryptocephalus sericeus Linné and Cryptocephalus globicollis Suffrian which the examined specimens were generally attributed to, when iden- tified. In our opinion ancient reports of C. globicollis from Amasya and Konya (Weise, 1884 ; 1901) must be assigned to the present species. Locus TYPICUS. Camlibeli Geçidi (Sivas, Turkey). DISTRIBUTION IN TURKEY. Amasya; Ankara; Artvin; Bilecik; Erzincan; Erzurum; Eskisehir; Kahramanmaras; Kars; Konya; Kiitahya; Sivas; Van. DERIVATIO NOMINIS. From the name of the ancient region of Northern Turkey. MATERIAL EXAMINED. Holotype (housed in Museo Civico di Storia Naturale, Milano): Anatolien, Sivas, Camlibeli Geçidi, 1500m, 30.7.1988, 1 5. Paratypes: Cerkes (30 Km E), Kursunlu, 27.7.1996, 1 d, (MZ); Cubuk, 3.8.1979, 1 3, (coll. Sassi); Artvin, Yusufeli, Altiparmak, 1400-1800m, 20/22.7.1983, 2 2 2, (MM and DE); Artvin, Yusufeli, Altiparmak, 1400-1800m, 4/8.8.1984, 2 9 ©, (coll. Sassi and MV); Obere Waldzone bei Altiparmak, 1500-1900m, 6/7.8.1978, 1 3, (DE); Artvin, Yusufeli, Ögdam, 1700m, 4.8.1984, 1 6, (DE) and 1 2, (USR); Bilecik, Osmaneli, 500m, 14.7.1972, 1 g, (coll. Sassi); 98 SASSI & KISMALI Umg. Pazaryeri, (Bilecik), 1.8.1984, 1 à, (coll. Sassi); Pülümür, Erzincan, 23/25.7.1988, 1 6, (DE); Akaysu, westl. Refahiye, 1800m, (Dumanle-dig), 18.VII.1985, 1 4 , (DE); Pass 2400m zw. Ispir u. Ovacik, 22.8.1972, 2 & 6, (DE); Pass bei Askale, westl. Erzurum, 26.7.1988, 1 6, (Muséum natio- nal d'Histoire naturelle, Paris); Erzurum, 2300-2500m, 14.7-2.8.77, 1 6, (HK); Erzurum, Nord Ispir, 1750/1900m, 11.6.1998, 1 3, (coll. Sassi); Kackar-Sudseite bei Hevek, (Meretet-yayla), 2000-2500m, 21.7.1983, 1 6, (DE); Eskisehir, Sögüt, 25.7.1988, 2 6 d, (HK); Konya, Sultan Mts, Cay, 15.8.1971, 1400-2000m, 2 2 ©, (DE); Bozkir, Hadim, 21.6.94, 1 4, (MZ); Manisa, Boz dag, 1500m, 7.1973, 1 3, (coll. Sassi); Manisa, Boz dag, 30.7.1972, 1 à , (coll. Sassi); Umg. Kütahya, 1 à, (coll. Sassi); Dumlupinar, 29.7.1997, 1 3, (coll. Kismali); Samsun, betw. Samsun and Kavak, 300m, 11.7.1976, 1 4, (USR); Sarigel (25 Km N), Bicakcilar, 13.7.1996, 1 dg, (coll. Sassi); Sivas, Camlibeli geg., 1500m; 30.7.1988, 2 2 ©, (coll. Sassi and HK); Sivas, Pagabahçe, 1250m, 15.7.1979, 1 ©, (coll. Kismali); Pass 2300 West Gevas, (Kuzgunkiran gec.), 13.7.1985, 1 5, (MV); Pass 2300 West Gevas, (Kuzgunkiran geç.), 23.7.1985, 2 6 à, (coll. Kismali and DE) and 1 ?, (DE); Tarmanli gegidi, Nordl Van, 29.7.1988, 1 2, (DE); Beytepe, 11.9.1977, 1 3, (coll. Sassi). DESCRIPTION OF MALES. Body 6.56 (+ 0.19) mm long by 3.99 (+ 0.12) mm, stout, Le convex, entirely metallic green. Antennae dull and blackish with the first segment shiny green. Frons smooth or slightly concave, closely and strongly punctured, with dense hairs. Antennae rather short, slender, second segment 1,3 times as long as wide, 6th - 11th seg- ment equal in lenght. Pronotum strongly convex, bulging, at base 1.34 as wide as long, forecorners with some scattered hairs; punctation strongly impressed and regularly distributed; interspaces between punctures almost shiny; lateral margins narrow, visible simultaneously only at base; not broadening toward hindcorners. Scutellum raised, as long as wide, slightly punctured, truncate at apex. Elytra 1.7 times as long as pronotum and 1.14 times as long as wide at humeri, squared, parallelsi- ded, convex, with more elevated postscutellar area, strongly and irregularly punctate; punc- tures on slope slightly smaller than in median part of disc; the intervals between the ely- tral punctation slightly convex and sparingly micropunctured; humeral tubercles promi- nent, lengthened; margins of elytra simple, barely visible from above. Epipleura with spar- se punctures. Pygidium regularly convex in middle, with deep and dense punctation and covered with rather dense withish hairs as well the abdomen and legs. Legs and ventrites metallic green. Anal sternite in males with a transverse depression; posterior margin of anal sterni- te straight or slightly notched. Fore basitarsi in males only slightly enlarged and rather short. Apex of aedeagus (figures 36-38) triangular, clearly sinuate; fraena of endophallus trian- gular. DESCRIPTION OF FEMALES. Body 7.70 (+ 0.39) mm long by 4.68 (+ 0.28) mm. Pronotum more transverse, at base 1.43 times as wide as long. Elytra are 1.8 times as long as prono- tum and 1.2 times as long as wide at humeri. Spermatheca (figure 46) sickle shaped; proxi- mal part swollen; apex fairly tapered; gland tubular, without constrictions; ductus not coi- led. TAXONOMIC REMARKS. The species 1s closely related to Cryptocephalus sericeus Linné from which it differs in generally more slender apex of aedeagus; in finer pronotal punctation The Cryptocephalinae of Turkey 99 and shiny intervals; in lateral margin of pronotum narrower and not enlarged toward hind corners; in lacking the bilobate tubercle above the depression of male anal sternite. It is also very similar to Cryptocephalus globicollis Suffrian from which it differs in stouter body outline, in generally more impressed punctures on pronotum, in lacking the rounded pit above the transverse depression on male anal sternite, in the shape of first scle- rite of internal sac (figures 39-42), that has a generally less pointed and less outwardly flexed apical denticle and the anterior median margin regularly curved, without the addi- tional denticle usually present in C. globicollis (figures 43-45). BIOLOGICAL NOTES. Collected on Astragalus and Centaurea, according to labels data. Cryptocephalus (Cryptocephalus) paradisiacus Weise, 1900 Cryptocephalus paradisiacus Weise, 1900: 276. Locus TYPICUS: Mardin. GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: recorded from the type locality only. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. TAXONOMIC REMARKS: Type series seems to be lost, but on the basis of the original description C. paradisiacus could be a junior synonym of the very variable Cryptocephalus cribratus Suffrian. Cryptocephalus (Cryptocephalus) parvulus Müller, 1776 Cryptocephalus parvulus Müller, 1776: 58. TERRA TYPICA: Denmark. SYNONYMS: Cryptocephalus nigrocoeruleus Goeze, 1777; Cryptocephalus flavilabris Fabricius, 1787; Cryptocephalus coeruleus Olivier, 1791. GENERAL DISTRIBUTION: Europe; Iran; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Bursa; Konya; Samsun. MATERIAL EXAMINED: Burdur, Aglasun, 15.7.1972, 1 spec. BIOLOGICAL NOTES: Meso-hygrophilous species, associated with the tree layer. Polyphagous, it has been recorded from Salicaceae, Corylaceae, Fagaceae and Betulaceae. SOURCES: Weise, 1901; Tomov & Gruev, 1975; Medvedev, 1975; Lopatin, 1977; Rapilly, 1980; Cryptocephalus (Cryptocephalus) pelleti Marseul, 1875 Cryptocephalus Pelleti Marseul, 1875: 183. TERRA TYPICA: Eastern Pyrenees. SYNONYM: Cryptocephalus androgyne Marseul, 1875. GENERAL DISTRIBUTION: France; Corse; Spain (Warchalowski, pers. comm.), Iran; Italy ? (as C. androgyne); Greece ? (as C. androgyne).The actual distribution of this species is at present poorly known, as it is quite recently removed from synonymy with C. cae- rulescens. According to Rapilly (1979) it is possible that C. caerulescens would be con- 100 SASSI & KISMALI fined to Northern and Central Europe, and reports from Mediterranean Region, Transcaucasus and Central Asia might have to be referred to the present species. DISTRIBUTION IN TURKEY: new to Turkey. MATERIAL EXAMINED: Artvin, Damar, Murgul, 1.8.1997, 1 spec., (JB); Bursa, Ulu Dag, 800m, 25.5.1981, 3 specs., (USR); Erzurum, Narman, Devrekani, 19.7.1979, 1 spec.; Kars, Göle, 1700- 1900m, 9.7.1987, 1 spec., (USR). BIOLOGICAL NOTES: Mesophilous or hygrophilous species, associated with the tree layer (Salix). SOURCES: Marseul, 1875; Apfelbeck, 1901; Berti & Rapilly, 1973; Lopatin, 1977; Rapilly, 1979; Warchalowski, 1991. Cryptocephalus (Cryptocephalus) peyroni Marseul, 1875 Cryptocephalus Peyroni Marseul, 1875: 169. Locus TYPICUS: Beirut (Lebanon). GENERAL DISTRIBUTION: Syria; Lebanon; Jordan; Israel; Turkey. DISTRIBUTION IN TURKEY: Artvin. MATERIAL EXAMINED: Hatay, Cevlik, Samandag, 23/26.4.1994, 4 specs., (JB) and (MZ); Hatay, Tranepinar, 22/23.4.1994, 1 specs., (JB). BIOLOGICAL NOTES: On Pistacia palaestina Boiss. according with Lopatin & Chikanutov 199%) SOURCES: Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997a, 1997b. Cryptocephalus (Cryptocephalus) praticola Weise, 1889 Cryptocephalus praticola Weise, 1889: 1. TERRA TYPICA: Caucasus. GENERAL DISTRIBUTION: Caucasus; Ukraine; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Erzurum; Artvin. MATERIAL EXAMINED: Kirklareli, Yildiz Mts., Demirköy, 500/900m, 25.6.1997, 1 spec.; Samsun, 3... 1993.3 SPECS. BIOLOGICAL NOTES: no data. TAXONOMIC REMARKS: A substantial revision of the species of the C. hypochaeridis com- plex is being printed. Because of the problems within these species, the real distribu- tion of the present taxon is scarcely known. SOURCES: Aslan & Ozbek, 1997a, 1997b. Cryptocephalus (Cryptocephalus) quadriguttatus Richter, 1820 Cryptocephalus quadriguttatus Richter, 1820: 12. TERRA TYPICA: not recorded. GENERAL DISTRIBUTION: Eastern Europe; Ukraine; Caucasus; Central Asia; Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: none. The Cryptocephalinae of Turkey 101 Figs 36-38: aedeagus, (scale: 1.61 mm). Figs 39-45: First sclerite of endophallus, (scale: 0.99 mm). 36-42: Cryptocephalus paphlagonius n. sp. (36-38 and 40: Sivas, holotypus; 39: Mts Kackar; 41: Bilecik; 42: Piiliimiir). 43-45: C. globicollis Suffrian (43-44: Alpes Maritimes, France; 45: Eastern Pyrenees). 102 SASSI & KISMALI BIOLOGICAL NOTES: Biology of this species is very poorly known. Burlini (1956) repor- ted Pinus and Alnus as possible host plants, also collected on Salix, Populus and Rosa according to Medvedev & Roginskaia, (1988). It could be defined a mesophilous spe- cies associated with the tree layer. SOURCES: Apfelbeck, 1916; Burlini, 1956; Ogul, 1970; Lopatin, 1977; Medvedev & Roginskaia, 1988; Warchalowski, 1991. Cryptocephalus (Cryptocephalus) rugicollis Olivier, 1791 Cryptocephalus rugicollis Olivier, 1791: 611. TERRA TYPICA: Var (France). SYNONYM: Cryptocephalus virgatus Suffrian, 1847. GENERAL DISTRIBUTION: Southern Europe; Jordan; Northern Africa; Turkey. DISTRIBUTION IN TURKEY: Amasya; Antalya; Aydin; Mugla; Izmir. MATERIAL EXAMINED: Aydin, Söke, 20m, 4.5.1991, 2 specs., (USR); Izmir, 1 spec.; Izmir, Cesme ilica, 13.4.1979, 1 spec.; Izmir, Efes, 3.5.1963, 1 spec.; Izmir, Yenisakran, 28.4.1984, 3 specs. (USR); Makri, Taurus, 6 specs.; Manisa, Manisa Mt., 5.1994, 1 spec., (JB). BIOLOGICAL NOTES: Xerophilous species, strictly associated with the Mediterranean belt. Oligophagous, it feeds on various genera of Asteraceae. SOURCES: Sahlberg, 1913; Kocher, 1958; Tomov & Gruev, 1975; Medvedev, 1975; Lopatin & Chikanutov, 1997; Aslan & al., 1977. Cryptocephalus (Cryptocephalus) ruguliventris Franz, 1949 Cryptocephalus hypochoeridis ssp. ruguliventris Franz, 1949: 185. TERRA TYPICA: Georgia. TAXONOMIC REMARKS: A substantial revision of the species of the C. hypochaeridis com- plex is being printed. Because of the problems within these species, the real distribu- tion of the present taxon is scarcely known. GENERAL DISTRIBUTION: Caucasus; Romania; Turkey. DISTRIBUTION IN TURKEY: Artvin (1000m); Rize (1200-1400m). MATERIAL EXAMINED: Artvin, 12.6.1973, 1 spec.; Artvin, 23.6.1990, 3 specs., (DE); Artvin, Artvin- Yusufeli, 500-1000m, 2 specs.; Artvin, Pass between Murgul and Arhavi, 1000-1200m, 6.8.1973, 1 specs., (DE); Artvin, Karckal-Otingöl, 600m 20.7.1987, 1 spec., (DE); Artvin, Karckal-Otingöl, 1800-2300m, 11.8.1978, 1 spec., (DE); Artvin, Savsat, 1300-1600m, 6.8.1970, 1 spec., (DE); Artvin, Yusufeli, Altiparmak, 1400-1800m., 20.7.1983, 5 specs., (DE); Erzurum, Kackar Mt., Heyek, 2000-2500m, 21.7.1983, 1 spec., (DE); Kars, 6.7.1975, 2 specs., (DE); betw. Artvin and Rize, Mt. Kackar 22.8.1973, 1 spec.; Artvin, Sarigöl, Bicakeilar, 13.7.1996 1 spec. (JB); Rize, Ikizdere, 1000-1500m, 25.6.1997, 2 specs. (CSAL). BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Tomov & Gruev, 1975; Tomov, 1984; Warchalowski, 1991. Cryptocephalus (Cryptocephalus) sericeus (Linné, 1758) Chrysomela sericea Linné, 1758: 374. TERRA TYPICA: not recorded. The Cryptocephalinae of Turkey 103 SYNONYM: Cryptocephalus robustus Suffrian, 1853; Cryptocephalus bidens Thomson, 1868. TAXONOMIC REMARKS: The status of the geographic forms attributed to this species is at present object of revision by one of us (DS). In this work we prefer to consider C. seri- ceus as a single widespread and variable taxon. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Siberia; Turkey. DISTRIBUTION IN TURKEY: Ankara; Erzurum; Kars (1800m). Some of the records might have to be attributed to C. paphlagonius n. sp. MATERIAL EXAMINED: Artvin, Savsat - Carn Gec., 1000/1500m., 26.6.97, 1 spec. (CSAL); Bozdag, 30.7.1973, 1 spec.; Bursa, Cagliyan, 10/14.7.1997, 1 spec. (JB); Cankiri (Kastamonu), Ilgaz- Dagh, 1800-2000m, 13.8.65, 1 spec., (DE); Izmir, Zeitingdag, 7.1973, 1 specs.; Kars, Ardahan, 6.7.75, 1 spec., (DE); Kars, Ardahan, Hanak, 1900-2100m., 7.8.1970, (DE); Kars, Sarikamis, 28.6.97, 1 spec. (CSAL); Kars, Sarikamis, 2000m, 3.8.1985 1 spec., (DE); Kars, Sarikamis, 2200m., 26.7.1983 e 16.8.1978 2 specs., (DE); Kirklareli, Kiyiköy, 3.6.1975, 2 specs.; Samsun, Samsun - Kavak, 300m, 11.7.1976, 1 spec.; Sivas, 15.7.1979, 1 spec. BIOLOGICAL NOTES: Meadow mesophilous species, polyphagous but clearly oriented to oligophagy, feeds on Asteraceae and locally on Ranunculaceae and Knautia. SOURCES: Ogul, 1970; Tomov & Gruev, 1975; Lopatin, 1977. Cryptocephalus (Cryptocephalus) sexpunctatus (Linné, 1758) Chrysomela 6-punctata Linné, 1758: 375. TERRA TYPICA: Europe. GENERAL DISTRIBUTION: Europe; Balkans; Ukraine; Siberia; Japan; Turkey. DISTRIBUTION IN TURKEY: Erzurum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Polyphagous and mesophilous, associated with the tree layer (Fagaceae, Betulaceae, Salicaceae; Rosaceae). SOURCES: Burlini, 1956; Mohr, 1977; Warchalowski, 1991; Aslan & Ozbek, 1998. Cryptocephalus (Cryptocephalus) signatifrons Suffrian, 1847 Cryptocephalus signatifrons Suffrian, 1847: 172. TERRA TYPICA: not defined. GENERAL DISTRIBUTION: Southern and Central Europe; Turkey. DISTRIBUTION IN TURKEY: Erzurum; Malatya. MATERIAL EXAMINED: Izmit, Kandira, 19.5.1973, 1 spec. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous but with tendency toward oligophagy, it has been prevalently collected on Corylus (Corylaceae) and secundarly on Quercus (Fagaceae). SOURCES: Burlini, 1956; Medvedev, 1970; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) surdus Rapilly, 1980 Cryptocephalus surdus Rapilly, 1980: 81. Locus TYPICUS: Hatam-Bak (Khoramabad, Iran). 104 SASSI & KISMALI GENERAL DISTRIBUTION: Syria; Jordan; Iran; Turkey. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: none. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Rapilly, 1980; Lopatin & Chikanutov, 1997. Cryptocephalus (Cryptocephalus) tappesi Marseul, 1868 Cryptocephalus tappesi Marseul, 1868: 206. Locus Typicus: Mt. Kulegh (Adana). SYNONYM: Cryptocephalus bidens Suffrian, 1860. GENERAL DISTRIBUTION: Greece (?); Syria; Lebanon; Turkey. DISTRIBUTION IN TURKEY: Taurus Mts. MATERIAL EXAMINED: Adana, Gaziantep, Nurdagi gecidi, 1200m, 19.6.1992, 1 spec., (MZ); rene 5.1994, 5 specs., (JB); Hatay, Cevlik, Samandag, 23.4.1994, 1 spec., (JB); Mersin, Camliyayla, 7.6.84, 1 spec.; Mersin, Camliyayla, 950-1400m, 1.6.92 1 spec.; Mersin, Namrun, 5:97, 2specs*. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Pic, 1904, 1941; Marseul, 1875; Weise, 1893; Lopatin & Chikanutov, 1997. Cryptocephalus (Cryptocephalus) trimaculatus Rossi, 1790 Cryptocephalus trimaculatus Rossi, 1790: 96. TERRA TYPICA: Tuscany (Italy). SYNONYM: Cryptocephalus salicis Fabricius, 1792. GENERAL DISTRIBUTION: Spain; France; Italy; Balkans, Transcaucasus; Syria; Jordan; Egypt; Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Agri, (1800m); Amasya; Ankara; Artvin; Bursa; Erciyas; Erzurum; Giimiishane; Konya; Mersin; Mugla. MATERIAL EXAMINED: Adana, Hasanbeyli, 30 Km E of Osmaniye, 17.7.1997, 1 spec., (JB); Adana, Pozanti, 20/29.6.1992, 1 spec., (JB); Antalya, Imrasan Gegidi, 1300m, 9.7.1993 and 22.6.1997, 3 specs.; Bilecik, Lalehan, Sögüt, 20.6.1978, 1 spec.; Eskisehir, Gunele, Sakari Ilica, 6/9.7.1997, 1 spec., (JB); Konya, Pinarbasi, 3.6.1970, 1 spec.; Mersin, 15 Km W Mersin, 1/5.7.1992, 2 specs., (JB); Mersin, Camliyayla, 1300m, 1.7.1993, 1 spec.; Mersin, Namrun, 35 Km NNW of Tarsus, 20.7.1997, 2 specs., (JB); Mersin, Tasucu, 3.7.93, 1 spec., (MZ); Mersin, Tasucu, 5.1994, 1 spec., (JB); Nevsehir, 28.6.1977, 1 spec.; Zonguldak, Safranbolu, 18.7.1979, 1 spec. BIOLOGICAL NOTES: Xerophilous species, associated with the tree layer; polyphagous, on Fagaceae, Rosaceae and Pistacia. SOURCES: Escherich, 1897; Bodemeyer, 1900; Ganglbauer, 1905; Tomov & Gruev, 1975; Mohr, 1977; Tomov, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) turcicus Suffrian, 1847 Cryptocephalus turcicus Suffrian, 1847: 173. Locus TYPICUS: Turkey. SYNONYM: Cryptocephalus pistaciae Suffrian, 1853. The Cryptocephalinae of Turkey 105 N: u Fig. 46: spermatheca. Figs 47-54: aedeagus. 46: Cryptocephalus paphlagonius n. sp. (Sivas), (scale: 1.00 mm); 47: C. sexpunctatus Linné (Suisse), (scale: 1.22 mm); 48: C. octopunctatus Scopoli (Northern Italy), (scale: 1.22 mm); 49: C. duplicatus Suffrian (Igel, Turkey), (scale: 1.18 mm); 50: C. concolor Suffrian (Mazendaran, Iran), (scale: 1.18 mm); 51: C. virens Suffrian (Slovenia), (scale: 1.18 mm); 52: C. violaceus Laicharting (Montenegro), (scale: 1.18 mm); 53: C. biledjekensis Pic (Izmir, Turkey), (scale: 0.76 mm); 54: C. cribratus Suffrian (Bolu, Turkey), (scale: 1.00 mm). Figures 47-52 show aedeagus in ventral view. Figures 53-54 show aedeagus in dorsal view. 106 SASSI & KISMALI GENERAL DISTRIBUTION: Southern Europe; Syria; Jordan; Turkey. DISTRIBUTION IN TURKEY: Ankara; Antalya; Balikesir; Bilecik; Bursa; Eskisehir; Istanbul; Mataraci; Trabzon. MATERIAL EXAMINED: Ankara, Kizilcahamam, 9.6.1978, 5 specs.; Burdur, Aglasun, 28.4.1972, 1 spec.; Cankiri, Cerkes, 22.6.1997, 2 specs. (CSAL); Isparta, Egridir, 27.4.1972, 2 specs.; Sivas, Camlibel, 25.4.1978, 1 spec. BIOLOGICAL NOTES: Xerophilous species, associated with the tree layer; probably oli- gophagous, it feeds on Fagaceae (Quercus), also collected on Corylus. SOURCES: Bodemeyer, 1900; Medvedev, 1970; Tomov & Gruev, 1975; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Cryptocephalus) virens Suffrian, 1847 Cryptocephalus virens Suffrian, 1847: 125. Locus TYPICUS: Kiew (Ukraine). TAXONOMIC REMARKS: Among the specimens from Suffrian collection (Martin-Luther Universitat, Halle), we could find one male labelled with the number 14372 (handw- ritten) that corresponds to the following data on Suffrian catalogue: Kiew, Hochhuth. Since data match the original description, we designated the specimen as lectotype of the species. GENERAL DISTRIBUTION: Central and Eastern Europe; Ukraine; Caucasus; Central Asia; Western Russia; Mongolia; China (?); Turkey. DISTRIBUTION IN TURKEY: Ankara, Erzurum. MATERIAL EXAMINED: Gümüshane, Kosedagi gec., 1400-1850m, 12.6.1998, 1 spec.; Kars, Sarikamis, 2000m, 7 and 11.6.1998, 11 specs.. BIOLOGICAL NOTES: Mesophilous species whose biology is poorly known, perhaps asso- ciated with the tree layer (Salicaceae), but most likely feeding on herbaceous plants (Asteraceae, Geranium, Onobrychis), similar to most related species (C. violaceus, C. sericeus and C. hypochaeridis complex). SOURCES: Escherich, 1897; Mohr, 1977; Gruev & Tomov, 1984; Medvedev & Roginskaia, 1988: Warchalowski, 1991; Aslan & Ozbek, 1998. Cryptocephalus (Cryptocephalus) volkovitshi Lopatin, 1976 Cryptocephalus volkovitshi Lopatin, 1976: 75. Locus TYPICUS: Arpachaystroy (Il'ichevsk distr., Armenia). GENERAL DISTRIBUTION: Armenia. We examined several specimens from Garni (Kotayk distr., Armenia, 7.7.1993) and Hatsavan (Abovian distr., Armenia, 14.7.1996). DISTRIBUTION IN TURKEY: Kars, Araxes valley, 18 Km NW from Idgir (Lopatin, pers. comm.). new to Turkey. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Collected on Atraphaxis sp. SOURCES: Lopatin, 1976. Cryptocephalus (Burlinius) bilineatus (Linné, 1767) The Cryptocephalinae of Turkey 107 Chrysomela bilineata Linné, 1767: 597. TERRA TYPICA: Europe. SYNONYMS: Pachybrachis armeniacus Faldermann, 1837; Cryptocephalus spitzyi Suffrian, 1848; Cryptocephalus partitus Jacoby, 1885. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Siberia; Korea; Japan; Turkey. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Meadow meso-xerophilous species. Polyphagous but prevalently associated with Asteraceae. Plumbaginaceae (Statice) have been reported by older authors, but that data need confirmation. SOURCES: Winkler, 1929; Ogul, 1970; Lopatin, 1977; Warchalowski, 1991. Cryptocephalus (Burlinius) chrysopus Gmelin, 1788 Cryptocephalus chrysopus Gmelin, 1788: 1713. SYNONYMS: Chrysomela biguttata Schaller, 1763 nec Scopoli, 1763; Cryptocephalus Hiibneri Fabricius, 1792. TERRA TYPICA: Europe. GENERAL DISTRIBUTION: Central and Southern Europe; Ukraine; Turkey. DISTRIBUTION IN TURKEY: Samsun. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous, it has been found on Rosaceae, Fagaceae, Corylaceae and Salicaceae. SOURCES: Medvedev, 1970; Ogul, 1970; Gruev & Tomov, 1984. Cryptocephalus (Burlinius) connexus Olivier, 1807 Cryptocephalus connexus Olivier, 1807: 836. Locus TYPICUS: Stralsund (Germany). GENERAL DISTRIBUTION: Southern Europe; Ukraine; Caucasus; Syria; Israel; Jordan; Iran; Central Asia; Turkey. DISTRIBUTION IN TURKEY: Ankara; Artvin; Bilecik; Bursa; Erzurum; Giresun; Isparta (1500m); Izmir; Samsun. MATERIAL EXAMINED: Adapazari, Hendek, 14.7.1972, 4 specs.; Adapazari, Sapanca, 15.7.1972, 1 spec.; Adiyamam, Desni, 29.7.1977, 1 spec.; Ankara, Dodurga, 13.7.1970, 2 specs.; Bolu, Diizce, 15.7.1979, 7 specs.; Bolu, Yigilca, 15.7.1979, 1 spec.; Bursa, 18.7.1972, 1 spec.; Bursa, Cagliyan, 10/14.7.1997, 2 specs., (JB); Eregli, 16.7.1979, 4 specs.; Gümüshane, 11.7.1978, 2 specs.; Hatay, Harbiye, 6.6.1972, 1 spec.; Kastamonu, Bozkurt, 20.7.1979, 1 spec.; Kastamonu, Catalzeytin, 20.7.1979, 2 spec.; Kastamonu, Inebolu, 19.7.1979, 5 specs.; Kastamonu, Küre, 19.7.1979, 1 spec.; Kastamonu, Tosya, 4.8.1979, 3 specs.; Kirsehir, Kaman, 18.8.1979, 1 spec.; Ordu, Kumru,13.7.1978, 5 specs.; Samsun, 11.7.1972, 2 specs.; Siirt, Sinan, 9.9.1977, 3 specs.; Tekirdag, 26.7.1973, 1 spec.; Trabzon, Arsin, 1.9.1973, 1 spec.; Zonguldak, 16.7.1979, 3 specs.; Zonguldak, Ulus. 17,7,1979 A.spec. BIOLOGICAL NOTES: Xerophilous species, probably associated with the tree layer (Ulmus) 108 SASSI & KISMALI and herbaceous plants as well (Statice) (Rapilly, 1979). Feeding on Pistacia lentiscus in Eastern Mediterranean (according to Lopatin & Chikanutov, 1997). SOURCES: Ogul, 1970; Lopatin, 1977; Rapilly, 1979; Lopatin, 1981; Tomov, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) elegantulus Gravenhorst, 1807 Cryptocephalus elegantulus Gravenhorst, 1807: 152. TERRA TYPICA: not recorded. SYNONYMS: Cryptocephalus tessulatus Germar, 1813; Cryptocephalus jucundus Faldermann, 1837. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Central Asia; Siberia; Mongolia; Korea; Turkey. DISTRIBUTION IN TURKEY: Giresun; Konya. MATERIAL EXAMINED: Ankara, Baglum, Sögüt, 9.7.1977, 3 specs.; Bursa, Cagliyan, 10/14.7.1997, 1 spec., (JB); Hakkari, Yiiksekova, 11.6.1975, 2 specs.; Van, Caldiran, 13.6.1975, 1 spec. BIOLOGICAL NOTES: Mesophilous species, predominantly associated with the herbaceous layer, in our opinion; polyphagous, it was collected on Asteraceae and Geraniaceae. Medvedev & Roginskaia (1988) also reported Artemisia and shrubs as possible host plants (Rosa, Prunus). SOURCES: Weise, 1901; Winkler, 1929; Burlini, 1956; Ogul, 1970; Lopatin, 1977; Tomov, 1984; Medvedev & Roginskaia, 1988; Warchalowski, 1991. Cryptocephalus (Burlinius) exiguus Schneider, 1792 Cryptocephalus exiguus Schneider, 1792: 204. SYNONYMS: Cryptocephalus wasastjernii Gyllenhal, 1827; Cryptocephalus amiculus Baly, 1873. TERRA TYPICA: not recorded. GENERAL DISTRIBUTION: Europe; Ukraine; Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Bursa. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous species. It seems to live both on herbaceous plans than on the tree layer. Polyphagous (Asteraceae, Salix, Betula, Populus Alnus). SOURCES: Burlini, 1956; Medvedev, 1970; Ogul, 1970; Medvedev & Roginskaia, 1988. Cryptocephalus (Burlinius) fulvus (Goeze,1777) Chrysomela Fulva Goeze, 1777: 321. TERRA TYPICA: not recorded. SYNONYMS: Cryptocephalus minutus Fabricius, 1792; Cryptocephalus ochraceus Stephens, 1829; Cryptocephalus signaticollis Suffrian, 1848; GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Jordan; Israel; Saudi Arabia; Western Siberia; Northern Africa; Turkey. DISTRIBUTION IN TURKEY: Antalya; Erzincan; Erzurum; Giimiishane; Izmir; Mersin. The Cryptocephalinae of Turkey 109 Figs 55-60: aedeagus. Fig. 61: dorsal pattern, (scale: 2.00 mm). 55-57: Cryptocephalus tappesi Marseul (Mersin, Namrun, Turkey), (scale: 1.54 mm); 58-61: Pachybrachis pentheri Ganglbauer (Kayseri, Turkey, Lectotypus), (scale of aedeagus: 0.55 mm). MATERIAL EXAMINED: Ankara, Ayas, 12.7.1978, 2 specs.; Izmir, Kozak, 21.6.1979, 1 spec.; Manisa, Salihli, 21.5.92, 2 specs., (SZ); Mersin, Silifke, Mamli, 12/15.5.1994, 1 spec., (JB); Mugla, Marmaris, 29.5.1996, 1 spec., (ZMH). BIOLOGICAL NOTES: Mesophilous species; it can be collected on meadows and on tree layer as well. Polyphagous; host plants belong to several families: Fabaceae, Asteraceae, Fagaceae, Salicaceae, Cistaceae and Labiatae. TAXONOMIC REMARKS: Burlini (1969) described Cryptocephalus fulvus subsp. schatz- mayri on specimens from Rhodos and Israel. Its range also comprises Jordan and Saudi Arabia, according to Lopatin & Chikanutov (1997). The taxon might occur in Middle Eastern part of the range of C. fulvus. In our opinion the status of this taxon needs to 110 SASSI & KISMALI _ be evaluated in the light of more recent systematic principles. Therefore we prefer pro- visionally to attribute the Turkish specimens to the nominal form. SOURCES: Sahlberg, 1913; Ogul, 1970; Burlini, 1969; Medvedev, 1975; Mohr, 1977; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) labiatus (Linné, 1761) Chrysomela labiata Linné, 1761: 169. TERRA TYPICA: Sweden. SYNONYM: Cryptocephalus digrammus Suffrian, 1848. GENERAL DISTRIBUTION: Europe; Ukraine; Siberia; Mongolia; Turkey. DISTRIBUTION IN TURKEY: Erzurum (2300m); Malatya; Ordu. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous species, associated with the tree layer. Polyphagous, it feeds on plants belonging to several families: Fagaceae, Corylaceae, Salicaceae, Rosaceae and Betulaceae. Sometimes it has been sighted on herbaceous plants. SOURCES: Ogul, 1970; Medvedev, 1970; Tomov, 1984; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) lederi Weise, 1889 Cryptocephalus Lederi Weise, 1889: 259. TERRA TYPICA: Araxes valley (Caucasus). SYNONYMS: Cryptocephalus oomorphus, Jacobson, 1895; Cryptocephalus ovulum, Jacobson, 1894 nec Suffrian, 1854: 30. GENERAL DISTRIBUTION: Transcaucausus, Turkmenistan; Iran; Iraq; Afghanistan; Turkey. DISTRIBUTION IN TURKEY: Balaban (as C. oomorphus). MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous or hygrophilous species. Labiatae (Mentha) and Salicaceae (Salix) have been reported as possible host plants (Berti & Rapilly, 1973; Rapilly, 1980). SOURCES: Berti & Rapilly, 1973; Medvedev, 1978; Rapilly, 1980; Lopatin, 1977, 1981, 1982, 1985; Warchalowski, 1999. TAXONOMIC REMARKS: We could not examine types of Weise’s collection, as they seems to be lost (Warchalowski, pers. comm.). On the basis of the original descriptions and comparing specimens of Cryptocephalus lederi from Afghanistan, (det. Lopatin) and specimens of C. oomorphus also from Afghanistan, (det. Medvedev), and material coming from Iran, we agree with the recent proposal of the synonymy: C. lederi Weise, 1889 = C. oomorphus Jacobson, 1895 (Warchalowski, 1999). Cryptocephalus (Burlinius) macellus Suffrian, 1860 Cryptocephalus macellus Suffrian, 1860: 53. TERRA TYPICA: Greece. GENERAL DISTRIBUTION: Europe; Syria; Israel; Iran; Egypt; Tunisia; Turkey. DISTRIBUTION IN TURKEY: Amasya; Antakya; Balikesir; Bilecik; Canakkale; Erzurum; The Cryptocephalinae of Turkey 134 Istanbul; Konya; Malatya; Mersin; Samsun. MATERIAL EXAMINED: Izmir, Kozak, 480m, 19.5.1992, 1 spec. (SZ). BIOLOGICAL NOTES: Xerophilous species, almost exclusively occurring in the Meditarranean belt. Polyphagous, it probably feeds on Fagaceae (Quercus) and Ericaceae. SOURCES: Weise, 1906b; Sahlberg, 1913; Medvedev, 1970; Medvedev, 1975; Tomov & Gruev, 1975; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) ocellatus Drapiez, 1819 Cryptocephalus ocellatus Drapiez, 1819: 201. TERRA TYPICA: Nuremberg (Germany). SYNONYMS: Cryptocephalus geminus Gyllenhall, 1827; Cryptocephalus ochropezus Suffrian, 1853; Cryptocephalus corsicus Pic, 1909; Cryptocephalus hummleri Pic, 1909. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Iran; Central Asia; Western Siberia; Turkey. DISTRIBUTION IN TURKEY: Amasya; Ankara; Artvin; Balikesir; Bilecik; Bolu; Bursa; Canakkale; Erzincan; Erzurum (2300m); Eskisehir; Gümüshane; Istanbul; Kastamonu; Mersin; Ordu; Rize; Samsun; Sivas. MATERIAL EXAMINED: Ankara, Lalahan, 7.6.1978, 2 specs.; Adiyaman, Nemrut Mts, 1.7.93, 2 specs., (MZ); Bolu, Abant, 1100-1400m, 21.6.1997, 2 specs., (CSAL); Corum, Koparan, 25.5.1978, 2 specs.; Diyarbakir, Lice, 12.6.1976 2 specs.; Hakkari, 13.6.1986, 1 spec.; Hakkari, Cukurca, 14.6.1976, 1 spec.; Kars, Karakurt, Aras river, 1400m, 17.6.1986, 1 spec.; Kiitahya, Orencik, 19.5.1971, 1 spec.; Mardin, 3.6.1976 and 14.6.1976, 2 specs.. BIOLOGICAL NOTES: Meso-hygrophilous species, associated with the tree layer (Salix, Betula, Corylus, Populus, Alnus). The reports on meadow vegetation (Mentha, Melissa) need confirmation. SOURCES: Weise, 1884 ; Escherich, 1897; Bodemeyer, 1900; Ogul, 1970; Berti & Rapilly, 1973; Medvedev, 1975; Tomov & Gruev, 1975; Lopatin, 1977; Rapilly, 1980; Tomov, 1984; Warchalowski, 1991; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) pusillus Fabricius, 1777 Cryptocephalus pusillus Fabricius, 1777: 221. Locus TYPICUS: Hamburg (Germany). SYNONYMS: Chrysomela minuta Herbst, 1783, Cryptocephalus verticalis Boheman, 1851; Cryptocephalus raphaelensis Gautier, 1861. GENERAL DISTRIBUTION: Europe; Ukraine; Caucasus; Turkey. DISTRIBUTION IN TURKEY: Turkey (without any explicit locality). MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Mesophilous or hemihygrophilous species, associated with the tree layer; polyphagous, on Salicaceae, Betulaceae and Corylaceae. SOURCES: Ogul, 1970; Warchalowski, 1991. Cryptocephalus (Burlinius) pygmaeus ssp. vittula Suffrian, 1848 Cryptocephalus vittula Suffrian, 1848: 63. 112 SAssı & KISMALI TERRA TYPICA: Europe. TAXONOMIC REMARKS: The status of this taxon is not yet defined, so the true distribu- tion is hard to state precisely. Some authors consider C. vittula as a simple synonym of C. pygmaeus. Some others treated C. vittula as a distinct species. We agree with the opi- nion of Warchalowski (1991) and consider C. vittula as a subspecies of C. pygmaeus occurring in the Eastern part of the range. GENERAL DISTRIBUTION: Italy; Balkans; Syria; Israel; Northern Africa (?); Turkey. DISTRIBUTION IN TURKEY: Afyon; Denizli; Izmir; Izmit. MATERIAL EXAMINED: Antalya, Fethiye, 6.6.1973, 3 specs.; Bursa, Cagliyan, 10-14.7.1997, 2 specs., (JB); Izmir, Bornova, 29.5.1979, 2 specs.; Izmir, Kinik Yayla köy, 24.6.1974, 1 spec.; Mersin, Silifke, 5.1967, 1 spec.; Mugla, Marmaris, 31.5.1966, 1 spec., (ZMH). BIOLOGICAL NOTES: Hemixerophilous species, associated with herbaceous plants. Oligophagous, feeds on various genera of Labiatae. SOURCES: Marseul, 1875; Sahlberg, 1913; Winkler, 1929; Burlini, 1956; Ogul, 1970; Tomov, 1984; Gruev & Tomov, 1984; Warchalowski, 1991; Lopatin & Chikanutov, 1997; Aslan & Özbek, 1997. Cryptocephalus (Burlinius) rufipes (Goeze, 1777) Chrysomela Rufipes Goeze, 1777: 321. TERRA TYPICA: not recorded. SYNONYM: Cryptocephalus gracilis Fabricius, 1792. GENERAL DISTRIBUTION: Europe; Northern Africa; Turkey. DISTRIBUTION IN TURKEY: Istanbul. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Hygrophilous species, associated with the tree layer. Polyphagous, it has been usually found on Salicaceae. SOURCES: Medvedev, 1970; Warchalowski, 1991. Cryptocephalus (Burlinius) strigosus Germar, 1824 Cryptocephalus strigosus Germar, 1824: 560. TERRA TYPICA: Dalmatia. GENERAL DISTRIBUTION: Southern Europe, Balkans; Turkey. DISTRIBUTION IN TURKEY: Erzurum. MATERIAL EXAMINED: none. BIOLOGICAL NOTES: Oligophagous species, feeding on herbaceous plants (Mentha, Thymus). SOURCES: Burlini, 1956; Warchalowski, 1991; Aslan & Ozbek, 1997. Cryptocephalus (Burlinius) sultani Pic, 1920 Cryptocephalus sultani Pic, 1920: 22. Locus TYPICUS: Eskisehir. TAXONOMIC REMARKS: Thanks to the kindness of M.me Nicole Berti we examined the The Cryptocephalinae of Turkey | 113 GA Figs 62-64 and 67-69: aedeagus. Figs 65-66: foretarsi, (scale: 0.99 mm). 62-64: Pachybrachis lati- collis (Suffrian) (Mardin, lectotypus of P. misellus (Weise), (scale: 0.66 mm); 65 and 67: P. mardi- nensis (Weise) (Aydinlar, Turkey); 66 and 68-69: P. limbatus (Ménétriés) (Amman, Jordan). Figures 67-68 show aedeagus in dorsal view, (scale: 0.76 mm). syntypes of the species from Pic collection: 1 male (designated as lectotype) with the following data: “type” (handwritten white label) / “Klein - Asien Eski-Chehir v. Bodemeyer” (printed, white label) / “type” (printed, red label). 1 female with the fol- lowing data: “Type” (handwritten white label) / “Asia minor Sultan Dagh v. Bodemeyer” (printed, white label) / “type” (printed, red label)/ “sultani Pic” (handwritten, white label). GENERAL DISTRIBUTION: Endemic species from Turkey. DISTRIBUTION IN TURKEY: Eskisehir; Konya. MATERIAL EXAMINED: Adiyaman, 26.7.1977, 1 female (uncertain identification). BIOLOGICAL NOTES: No data were found in the literature. 11.4 SASSI & KISMALI Cryptocephalus (Burlinius) tschimganensis Weise, 1894 Cryptocephalus tschimganensis, Weise, 1894: 68. TERRA TYPICA: Chimgan (Tien Shan, Uzbekistan). GENERAL DISTRIBUTION: Central Asia; Turkey. DISTRIBUTION IN TURKEY: Baba Dagh; Izmir. MATERIAL EXAMINED: Karpuzda, 29.7.1977, 1 female (uncertain identification). BIOLOGICAL NOTES: Feeds on Glycirrhiza, according to Lopatin (1977), dubiously on Salix, Populus and Acer according to Medvedev & Roginskaia (1988). SOURCES: Weise, 1894a; Sahlberg, 1913; Lopatin, 1977; Medvedev & Roginskaia, 1988. Cryptocephalus (Burlinius) tshorumae Tomov, 1984 Cryptocephalus (Burlinius) tshorumae Tomov, 1984: 376. Locus TyPIcus: Bogazkale (Corum, Turkey). GENERAL DISTRIBUTION: Endemic species of Turkey. DISTRIBUTION IN TURKEY: reported from type locality only. MATERIAL EXAMINED: Bolu, 30.7.1981, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. Cryptocephalus (Burlinius) variceps Weise, 1884 Cryptocephalus variceps Weise, 1884 : 161. Locus Typicus: Istanbul. SYNONYM: Cryptocephalus karsantianus Pic, 1914 n. syn. TAXONOMIC REMARKS: Medvedev & Samoderzhenkov (1989) downgraded the taxon to a subspecies of C. exiguus, according to the quite identical shape of aedeagus. On the contrary we prefer to keep the taxa apart at species level until more information about range of distribution becomes available. We examined the type series of C. variceps [(4 specimens, labelled “Turcia” and “Merk!” (handwritten, white labels), one of them, a male, designated as lectotype by Lopatin (1989)]. Morphological differences with C. exiguus seem to be more definite compared with what Medvedev et Samoderzhenkov had reported in their work. As a matter of fact, C. variceps differs in stouter body outli- ne, more transverse pronotum, in lack of the typical lengthened punctation, in lighter coloration of legs, in dorsal process of aedeagus subequal to ventral ones (distinctly shorter in C. exiguus). Besides, we designated the type specimen of Cryptocephalus karsantianus Pic. It is a male, labelled as follows: ‘“? Karsanti” (handwritten, white label) / “type” (handwrit- ten, white label) / “type” (printed, red label) / “karsantianus Pic” (handwritten, white label) / “karsantianus Pic” (handwritten, blue label). In our opinion it is identical to the present species, so we propose the new synonymy: Cryptocephalus variceps Weise, 1884 = Cryptocephalus karsantianus Pic, 1914. GENERAL DISTRIBUTION: Balkans; Caucasus; Iran; Turkey. DISTRIBUTION IN TURKEY: Adana (as C. karsantianus); Istanbul; Izmir. The Cryptocephalinae of Turkey | 115 MATERIAL EXAMINED: Adapazari, Hendek, 14.7.1972, 1 spec.; Bursa, Iznik, 27.5.1967, 1 spec. BIOLOGICAL NOTES: No data were found in the literature. SOURCES: Weise, 1884; Berti & Rapilly, 1973; Tomov, 1984; Gruev & Tomov, 1984; Aslan & Ozbek, 1997. QUESTIONABLE OR ERRONEOUS RECORDS Cryptocephalus (Asionus) pavloskii Lopatin, 1956. Oglobin & Medvedev (1956) descri- bed C. imitator on specimens from Talas, (Kalca) without any further specification. In the Zoological Record (1956: 336) the locality was reported as being situated in Turkey, but in fact itis NW Kirghizistan. Afterwards (Lopatin, 1963) the species was synonymyzed to C. pavloskii. Cryptocephalus (Asionus) reitteri Weise, 1882. It has been reported by Warchalowski (1974,1991) who erroneously interpreted an ambiguous datum of Burlini (1956) (Warchalowski, personal communication). Cryptocephalus (Lamellosus) laevicollis Gebler, 1830. Older reports from Turkey (Weise, Bodemeyer) preceded the description of the very close C. angorensis Pic, 1908. It has also been reported by Tomov & Gruev (1975) from Amasya and Corum, but afterwards the record was attributed to C. angorensis (Tomov, 1979). Therefore the presence of this spe- cies in Turkey seems most unlikely. Cryptocephalus (Cryptocephalus) concolor Suffrian, 1847. It has been reported for the region by several authors (Bodemeyer, 1900; Breit, 1918; Berti & Rapilly, 1973; Warchalowski, 1974; Tomov & Gruev., 1975; Tomov, 1984; Aslan & Ozbek, 1997b) but according to Lopatin (1984) the species does not reach the territory under study and the reports must be attributed to C. duplicatus Suffrian, 1847. Cryptocephalus (Cryptocephalus) globicollis Suffrian, 1847. It has been reported by Weise (1884) dubiously from Amasya and afterwards (1901) from Konya. In our opinion these records must be ascribed to the closely related Cryptocephalus paphlagonius n. sp. Cryptocephalus (Cryptocephalus) violaceus Laicharting, 1781. All Turkish male specimens labelled as C. violaceus in the examined collection were in fact C. duplicatus. Females are pratically indistinguishable from C. virens and differ from C. duplicatus only in faint chro- matic differences. In spite of the indications of some authors (Breit, 1918; Burlini, 1956; Aslan & Ozbek, 1997b) the presence of this species in Turkey must be considered uncer- tain, in our opinion. Aslan & Ozbek (1997; 1998) reported from Turkey the following species as belon- ging to the Turkish fauna; on the contrary they have never been recorded in the territory under study: Cryptocephalus corinthius Pic, 1914; Cryptocephalus creticus Suffrian, 1847; Cryptocephalus inaequalis Fairmaire, 1902; Pachybrachis flexuosus, (Weise, 1882). 116 SASSI & KISMALI Besides, the following taxa are also reported by Aslan & Ozbek (1997); they were all described as chromatic varieties of quite variable species, therefore they have to be consi- dered synonyms of the respective nominal forms, in our opinion: Cryptocephalus bipunctatus var. thomsoni Weise, 1882; Cryptocephalus ilicis var. oblitteratipennis Pic, 1900; Cryptocephalus ilicis var. prinkipensis Pic, 1908; Cryptocephalus pygmaeus var. lugubris Demaison, 1904; Cryptocephalus pygmaeus var. orientalis Weise, 1882; Cryptocephalus tappesi var. disconiger Pic, 1904. KEY TO GENERA 1 Scutellum invisible from above; eyes entire; body small, cylindrical, dorsally with short hairs arr ee te el ae gg: Lele gst Ml LOT, I a AS Stylosomus Suffrian, 1848 af PS O ie N e E De eee ek Pe UID LG PR EL SL OA 2 2 Pronotum narrowly marginate, not closely fitted to base of elytra; elytral base thickened . . . .3 - Pronotum not marginate, closely fitted to base of elytra; elytral base not thickened ........ 4 3 Pronotal surface with distinct pubescence; scutellum flat at apex . . . . Thelyterotarsus Weise, 1882 (this genus is not yet recorded in the territory under study, but some species have been reported from Caucasus, Iran, Jordan and Israel, so its presence is possible in northeastern or southern regions) - Pronotal surface glabrous; scutellum elevated posterad ....... Pachybrachis Chevrolat, 1837 4 Antenna long, slender, preapical segments at least 1 1/2 times as long as wide ............. LIGA RE LAN ARE AT SUR VENTES dubita, ne & Cryptocephalus Müller, 1764 - Antenna short, preapical segments about as wide as long; body small (length 1.7-2.0 mm) ... sa let eRe RO, SP, ORESTE OE Melixanthus Suffrian, 1854 (the genus Melixanthus is not yet recorded in Turkey, but two species occur in neighbouring ter- ritories, so their presence is possible in the area under study. They are: M. brunnicollis (Suffrian, 1857): abdomen reddish brown, elytra and pronotum ochraceous with darker markings. Egypt, Israel. M. granularis (Suffrian, 1857) (= M. jordanicus Lopatin, 1979): abdomen and upper part enti- rely yellow. Caucasus, Jordan, Saudi Arabia, Israel). KEY TO TURKISH SPECIES OF GENUS STYLOSOMUS 1 Body reddish-brown to blackish; elytra irregularly punctate; tarsi distinctly shorter than tibiae (nolnertegordedandhesezioniz i Lab Lar ie ant RT Rs subg. Microstylus Rey - Body yellow or ochraceous; elytral punctures arranged in regular rows; tarsi lengthened, slen- der Ae Ones MIMBO MONON SE), aie e PIRA songe RE LE oe 2 2 Body unicolored yellow; elytral intervals with short, erect hairs; length 1.6 - 2.0 mm .../flavus - Vertex, base of elytra and suture generally darkened; elytral interval with long, closely fitting, SSG ey Ale Mae! tee eee REA RR A re RIS ee Seas beet VE 3 3 Pronotum deeply and regularly punctate; interstices on elytra flat, larger than the diameter of Poot utent ZI LIE EU LA Ee a e. tamaricis - Punctures on pronotum slightly smaller and sparser; interstices on elytra not larger than the dia- mer ofimmennes one? Odie ee N subelongatus The Cryptocephalinae of Turkey | TIR KEY TO TURKISH SPECIES OF GENUS PACHYBRACHIS 1 Pronotum entirely ochraceus with or without undefined reddish pattern ................. 2 i. At least pronpum with. avell developed lack pattern O. este zent nah data i 4 2 Elytral punctation arranged in regular, curved rows, additional punctures only in scutellar area; lengthy, 3.0 = RO mm. i Acie A cee e LENS RSR CT te a glycyrrhizae - #lytral punctures not arranged in regula Toys siete 3 3 Smaller species; head with well defined black dark pattern; elytral punctures brown at bottom as well as humeral spot; aedeagus with a simple triangular apex; length 3.4 - 3.8 mm ....... rg rere re els re Ae Oe ee anatolicus - Larger species; frons with darker pattern reduced to a reddish median line; dorsal surface total- ly ohraceous; elytral punctures very deeply impressed; aedeagus tridentate; length 3.0 to 5.0 mm idiots azadunent seo Mira attenzioni Wei STE US Zn 3 nigropunctatus 4 Pronotum with M - shaped black pattern; elytra yellow, only a little black spot on humeri, punc- tation partly in regular rows, punctures blackish at bottom; length 2.9 - 3.7 mm . .scripticollis Vi. Elytra and pronotuny with, well markedidler patterns deo, 5 I. Sides af-mesosternum. yellow. o. WIR A 221 oW. spot. sie anne sanre AT bernie hind 6 Ur Sile OF mesasternum completely Black nun NE PR ES 13 6 Length less than 3.2 mm, punctation on pronotum very dense and regular; elytral black pattern consists of a longitudinal stripe extending from base to clivus and three black spots on side (stri- pe and spots sometimes reduced); aedeagus as in figures 58-60; dorsal pattern as in figure 61 . i air sario, Anto re a iii ita pentheri - Length generally more than 3.2 mm; punctures on pronotum deeper and sparser .......... 7 7. Elytra mostly black, light pattern reduced to little yellow spots in relief; aedeagus as in figures 62-64: leneth,. 3.0.» 3:0: na... è Seta ice ian er er hte iano o laticollis «y Light pattern on-eljtra plane and darren fire earn nt ri à 8 8 Elytra yellow with suture, apical and lateral margins black; black pattern on disk forming seve- ral longitudinal furrows, particularly on sides, leaving intervals in relief; pronotum yellow with M-shaped black pattern; ventrites black; length 4.0 - 4.3 mm .................. adaliensis - Black pattern on elytra variable, but never forming longitudinal furrows ................ 9 9 Elytra yellow with black humeral spot (sometimes a short irregular stripe near lateral margin and a sign of postscutellar spot black as well); length 3.3-4.0mm.............. humeralis «| „Black pattern on.elvira more developed, ssc; ira elia den 10 10 Last abdominal sternite black; pygidium black or sometimes with little yellow spots; femora often with black line on dorsal surface; metatibiae sometimes darkened; length 3.0 - 4.0 mm . mare ere Feat Desire ee at ere tt Rl, Lae Ree ae hieroglyphicus =, : Pygidium.and last abdominal sternite with yellow spots. (tows esha basco tees a: 11 11 Smaller species (length of male less than 3.5 mm.); aedeagus with a short but distinct tooth on apex: length 2.8 - 3.5 mM . à «raw Pts aaa earl aan oat ead instabilis «1 Larger species (length of male more than. 3:3 ml. zu ai 12 12 Legs entirely yellow; punctures of postscutellar area sparser and partly arranged in subregular rows: lensth 40-45 mi SE ee albicans - Legs partly darkened; punctation on postscutellar area closer and completely confused; length 30-40 WM iii ea i a ee scriptidorsum 13 Body smaller; punctures on pronotum dense and more delicate; elytra mostly black with poste- rior half of suture, last interstice (posterior to humerus), apical and anterior margins and some sparse spots on disk yellow; length 2.4 - 3.4 mm ................fimbriolatus spp. mendax - Body larger, length more than 3.3 mm; punctures on pronotum generally deeper and sparser; posterior half of suture and last interstice at least partly black .. 0 ONE 14 118 SASSI & KISMALI 14 Elytra mostly black, light pattern completely missing or reduced to little yellow spots in relief - Yellow pattern on elytra plane and larger; elytra bene) yellow with three outer spots and a longitudinal black band extending in the middle of disk, from base to slope, often partly fused with sutural stripe; black spots and bands often coalescent; in the darkest forms, dorsal surface ANNE sota baie RAS Wee ern re LARP ORS E Ee as cg RA IAT ll i. elo partcmanmissnme: cluinacomplovely black. Aa N Ure. ees EN erw ans. (generally female specs. of P. limbatus, P. tessellatus, P. mardinensis and P. laticollis, there is at present no valid character for a correct identification) gem: OTTO SEN ee I ER EP EN on oe gta a 16 16 Apex of elytra with a yellow fimbria extending anteriorly; pronotum black with yellow lateral and anterior margins, additional light pattern consist of a longitudinal median line and two late- ral triangular spots fused with anterior margin, and two curved spots near base; elytra black with (OO CIS EDO ee ea ah a Para et Laie OA al - Apex of elytra yellow but without fimbria extending anteriorly; pronotum black with lateral and antemori mann veloweswithout additonalspots “LY Ara. ag talkie AR LRU. 19 17 Apical 1/3 of commissure generally yellow; meso- and metatibia almost completely or com- pletely black; punctures on disk of pronotum sparser and deeper; aedeagus notched terminally; NEU EC ORI EUR PRE ELE APR PE ER Ce PL PET PL DES SPORE PIERI AT: tessellatus ssp. orientalis - Yellow apical margin of elytra not extending anteriorly along commissure; meso- and metatibia yelowor palsy darkencd: aedeagus poimted ternmanally 2.722. 2 EN A ose le. 18 18 Body outline slender; smaller species (length of males about 3 mm); punctures on pronotum finer and more regularly distributed; epimera of mesothorax in the examined specs. with a little yellow spot; length 3.0 - 3.6 mm. x Medals | . .laticollis - Body outline stouter, larger species doi at en bal 3. 5 ana Sunctiires on PIREO dee- per and sparser; epimera of mesothorax completely black; length 3.5 - 4.0 mm ....... picus 19 Median and hind legs entirely black, with a withish spot on apex of femora ............. 20 - Tibiae, tarsi and base of femora partly yellowish; length 3.0 - 3.6 mm ............ laticollis 20 In male third tarsal segment not enlarged (figure 65); posthumeral spots on last and second-last interval partly fused together, the outer one more extended posterad; apex of aedeagus (figure VEIL CT gl IE ZUI PORTI CMGI A) a aa tae RCE ee RU, mardinensis - In male third tarsal segment fairly enlarged (figure 66); posthumeral yellow spot on last inter- stice small, sometimes obliteated, anyway not extending posterad more than the one on the second-last interstice; apex of aedeagus ( figures 68-69) pointed and turned upward; length 3.0 PA CDT ee TORI A ta rr a bec orga ob rca oe RO RE BOUM (UP € limbatus 21 Lateral interstice of elytra entirely yellow; hind tibiae darkened toward apex ............ 22 - Lateral interstice of elytra partly or completely black; hind tibiae entirely black; apical 1/4 of CW ica Complete selon lowe ore Sm, RR UE lees anes velarum 22 Apical 1/4 of elytra completely yellow; length 3.0 - 3.5 mm .................. bodemeyeri =~ ‘Only apical’1/6 of elytra completely yellow: tenet 32,3. 85mm N. sinuatus KEY TO TURKISH SPECIES OF GENUS CRYPTOCEPHALUS Lin lata comm letelyor pantianezilaria a Punetde.: 2. survenant i ale AEA wes 2 ir Blu Baal ae en ee teed ti ds 36 Zabel enne alli ee rasche aa à + 3 6, Dotsal surfdce alice Crary, MIT age wuss sodas rads Er ale + 32 ie eI eR OSS CTO I ee ar in ia oe ae + The Cryptocephalinae of Turkey | DS Ne) 10 13 14 17 Leñeth more than Sato FEIERTEN. EMEA 22 Aedeagus (figures 1-3) terminally ended into simple triangular apex; pronotum black, with yel- low anterior and lateral border, strongly and closely punctured; elytra yellow with a blackish stripe from humeri to clivus; suture margined with a black band; puncturation strong, regular, but rows are broken in posterolateral part of disc; interstices distinctly micropunctured; length 2.8-3:5 mini A LE ATMEN RETE SRO, BEI curda Apex of aedeagus with two ventral and generally one dorsal median process; rows of punctures conipletely feeularonitelyiran DEZENT III O) IRENE RI IPA EIERN 5 Pronotwm deeply punctureckorsiniate:surtacedul) on silly. ly Me IA 6 Pronotum impunctate or very finely and sparingly punctate, surface lustrous ............. 8 Pronotum withidense lone striations; surface appears lg E ee eee N, 7 Pronotal punctures rounded or slightly lengthened; length 1.7-2.3 mm .......... elegantulus Pronotum black; elytra black, with a median yellow band from humeri to clivus; length 25-25 im eg 4 Los We, Le ee ER ARE RA RE bilineatus Pronotum and elytra uniformly black or piceous; length 2.0 - 2.5 mm ............. exiguus Pronotum black or brownish black, at most lateral and anterior margins yellow ........... 9 Pronotum pale; vello or Teddies att I OR RIE REI), Rts eae 16 Elytra yellow with humeral blackish spot sometimes extended in a longitudinal stripe, suture Black. 24,9: Va Fe API, AIR eS PIE SEIEN. 10 Elytra completelsor mostly lack inc?) ee, EINE REN TIRES SIEB 11 Puncture of pronotum visible only with great magnification; anterior and lateral margins yel- low, light pattern gradually shading off to black of disc; supraorbital grooves deeply impressed, and widely separated from internal ocular ridges; length 2.5-3.0mm............ connexus Pronotum with delicate but easily seen punctures, anterior and lateral margins yellow as above, but bound with black of disc sharper; supraorbital grooves short, scarcely separated from ocu- laredses; length 2.20 Pa ture CP RS PO vittula Elytra black with yellow pattenti.. 4 705 RO IR EE EER oa 12 Elytra entirely black or piceows Hy. 2 AT ARI OI RIE IT 18 Elytra black with vellow-spot on apex; length 2:2 »2. 3m: men ein chrysopus Elytra black with a transverse yellow band near base and a lateral yellow stripe in the anterior half, partly extended to-epipleura; length 2-3 mia, FE oe strigosus Head black with a yellow rounded or elliptic spot upper eye-lobes .................... 14 Head without such spot’ Las € i+ Whee IE, E RS enr 15 Spots upper eye-lobes at same level as remainder of frons; antennae robust, preapical segments about 1.5 times as long as wide; dorsal process of aedeagus (figures 4-5) distinctly shorter than lateral ones; length'2.8:.4 Oi Fag na Je FE PERSE SII lederi Spots upper eye-lobes shiny and slightly elevated; antennae slender, subapical segments more than 2 times as long as wide; dorsal process of aedeagus ( figures 6-7) as long as lateral ones; length’ 2.5 - 3.3 mi ois ed PE eee Fak AA ea ocellatus Frons entirely or almost entirely yellow; clypeus and legs yellow as well; length 2.0mm .... ER IE ORE MAS EN variceps Frons blackish; clypeus and genal spaces yellow; femora partly darkened; length 2.2 - 3.0 mm Whee ace i E MR ORI OR SE labiatus Supraorbital grooves deeply impressed and widely separated from internal ocular edge; elytral rows fairly impressed, punctures clearly visible on apical clivus; apex of aedeagus trilobate, dor- sal process rounded? length 2.322: Fam) enc. Shop ee ee es Supraorbital grooves short and scarcely separated from ocular edge .................. 17 Pronotum longer and more convex; elytra often with indistinct ochraceous spots on humerus 120 SASSI & KISMALI and clivus, sometimes fused together to form a longitudinal band; rows impressed, lateral inter- vals convex; aedeagus (figures 8-9) without dorsal median process; length 2.1 - 3.2 mm ..... RE EEE AVA Ra RE ed ero. pete De‘ tschimganensis - Pronotum more transverse and less convex; apex of aedeagus trilobate; elytral rows weakened uso pie anc eee NE ee Pie re eat ua mme ls a | 18 iS Persalsumace yellow, iiidersideiblack an typical SPECS. iis daw ue. ane TUNEL A 19 - Pronotum reddish elytra with variable brown to black pattern, in extremely dark form almost whole disc is brown or black except epipleura and anterior part of lateral margins ........ 20 19 Pronotum short; elytral punctation more delicate, intervals flat, suture yellow, sometimes an indi- stinct ochraceus oblique stripe along each side of elytral disc; dorsal process of aedeagus poin- [cereale 2 22 Sr SERE ere tad ba earn macellus - Pronotum less transverse; elytral punctation more impressed, intervals convex, suture black; dorsal process of aedeagus rounded terminally; length 2.3 - 2.6 mm ............... sultani 20 Elytra black, yellow stripe on lateral margins not extended to posterior half of elytra, its bound with black of disk sharper; apex of elytra generally without yellow markings; length 2.5 - 3 RIA he ee nae ee ee EDER a the Ae Auto vor ad wha hed rufipes - Elytra brown to black, yellow stripe on lateral margins gradually shading off to black of disk and generally extended to posterior half of elytra; apex of elytra with yellow markings; mesoe- (Re D TOUR PE RE ERE RE E OE LEEREN rs hs PT 21 21 Pronotum impunctate; dorsal process of aedeagus (figures 10-11) rounded apically; length 2.5 RINO Ayer Remar” E ME Re lan all ious dares. ode pusillus - Pronotum minutely punctured; dorsal process of aedeagus (figures 12-13) pointed apically; length CAZIONE Pie es ee i Ae eee dolo enn Tin) aati ee, Lem Cie, ald vi tshorumae raie ce hello mac nn nant Gow: an Sle. nb aw en, 23 + Ely yellom-orvedewathnion pattern eigen Ooi te Male in Meli. 27 23 Pronotum black; elytra black with broad yellow spot on apex; length 4.5 - 6.0 mm... .biguttatus -¥) Wellompattenvextended so-anieniompartotelytta Le Lit AU ad de RL | 24 24 Base of elytra black; pronotum distinctly punctate; length 3.5 - 4.5 mm ............ moraei - Yellow elytral pattern comprises a basal band; pronotum impunctate .................. 25 25 Yellow bands at base of elytra include entirely the scutellum; rows of punctures deeper, clearly Cisiple ones lenein 354mm: our LL aus snusaios at nn «i octacosmus - Yellow bands at base of elytra do not reach the scutellum at their median side; rows of punctu- Repeats On ae PEUR EN some ere IVO DET. 26 26 Body outline stout and oval; elytra clearly converging towards apex; rows of punctures very fine; light spatter on elytra-ochraceons; leneth 3:5-'4:5 mm: 2, 0 00. 2 dee crassus - Body outline slimmer; elytral sides parallel, elytral light pattern whitish, often reduced or obli- Lena Ze A a dae vcs Hi amis dede eho. ok anticus iar eoayentwtely yollows lene 2.4475 mies, es nalen sa CO ee eal surdus albori dee ee issu ack hs Oo ee alee vo eh men CLL Ri II, 28 28 Pronotum with dark M-shaped pattern; elytra with sinuate, transverse bands; interrow spaces minnicly punctate; lenti: 33-00 mm. oo ea a a had da à va dun Hee ne ope eo so ows „ulmenifer - Elytra without sinuats)transverse alten ur santa ann ben copia dti. . 29 20 Franotum red, with dark spots; lenety6:0= 7.7 MM, iii gloriosus nny ecu ansia leanzttà el bel dà Lun ine ules ery ln. 30 30 Lateral margins of pronotum entirely visible from above, anterior angles often with some scat- tered hairs; elytra yellow or reddish with three black spots (1 humeral + 2 on apical clivus); lan, fe 3 ee oe ii Whoo, bau pente apo Job, imperialis - Lateral margins of pronotum not entirely visible from above, anterior angles bare ........ al The Cryptocephalinae of Turkey 121 DI 32 33 36 37 40 41 Elytra red, with three black spots (1 humeral + 2 on apical clivus); length 5.0 - 7.0 mm ...... er PEER EC Or et e RL N trimaculatus Elytra red, with variable black pattern, but never 1 + 2 black spots: 1 + 1 or only 1 humeral spot (sometimes elongate toward clivus); paler specs. have elytra completely reddish; length 4.0 - ON, RON le SME A la eRe BEE ER SRE. A ISAIA IDO bipunctatus Pronotum black with minute punctation, anterior and lateral margins yellow; elytra metallic blue; rows of elytral punctures weakly impressed, barely visible on apical clivus; legs completely yel- low; aededous' as nTigires14216 enah : IH THE Louis MR a peyroni Nopeucha. combination ol.charaster. ad au a ra bt ee; 33 Pronotum with dense punctation, larger and deeper on flanks; lateral margins of pronotum nar- row, visible from above only posteriorly; clipeus and frons (at least on apex of eyes) with yel- low pattern: lensth 3:9 +43. mm sa Merl a I HE ALERT ario Punctures on sides of pronotum not larger than in central part of disc; lateral margins of prono- tum wider, entirely visible from above; only clipeus and genae yellowish .............. 34 Pronotal punctures more impressed, punctures near anterior margin as deep as those on disc; dorsal surface entirely metallic in both sexes; length 3.0 - 4.0 mm ............... parvulus Pronotal punctures shallower, punctures near anterior margin less impressed than those on disc; elytral/surface-entirely metallic or mostly yellow inafemale wae He Een 35 Ventral surface of aedeagus (figure 17) with two longitudinal oblique furrows on apex; female with elytra mostly yellowelensıh 4.2 iR HR Literie ARE bicolor Ventral surface of aedeagus (figure 18) without longitudinal oblique furrows; upper part enti- rely metallicin both sexessdencth 4.0 1 A Ha PIE N Bi pelleti Protibiae in male specs. expanded in a triangular tooth on apex; apex of aedeagus strongly wide- ned, obtuse, spoon -shaped, with a median ventral lobe shortly surpassing the apical rim; late- ral margins of pronotum slender, not entirely visible from above; elytra red, with two spots near base and a transverse band on clivus black; underside, head and legs black, sometimes protibiae paler: subg. Lamellosus (only male specs. identifiable for certain). ................... 37 Not such a combination of charaéters ara a pi BAS LISTE et ah, 38 Triangular tooth on apex of protibiae smaller; lateral margins of apex of aedeagus (figure 19) not or scarcely bent ventrally; length 4.0 - 4.5 mm (uncertain presence in the territory under study)... 4. te Ree ed hed ans ASSES Be METIERS Ot ea RES Re: laevicollis Triangular tooth on apex of protibiae larger; lateral margins of apex of aedeagus (figure 20) clearly bent ventrally; lenethi4,7 = Sa Sul eR eee angorensis Ath tarsal segment slimmer, at least one half its length projecting out of lobes of 3rd segment; elytra more or less hairy, in particular on sides and clivus: subg. Asionus, pars ........... 39 4th tarsal segment stouter and shorter u. oF eb SEL REL HUM a AH Metallic blue; anterior and lateral margins of pronotum yellow in male, elytra with yellow band on anterior half of lateral sides; apex of elytra with yellow spot; legs darkened, apex of femora with pale spot: length 40-30 mm HH, Du he A BIN apicalis Dorsally not metallic, yellow orsed with dark pattern are 40 Body stouter (length / width ratio less than 1.8); pronotal punctation coarse and dense; prono- tum black or brown with-vellow:pattettvvo.i pri dy leu Gy Bea RLS ee cu 4] Body slender, cylindrical (length / width ratio = 1.9 or greater), pronotum yellow with little black spots; length 5.0 mm or mare «2 2 Oi Sad ebbe A da 42 Pronotum mostly brownish, with yellow anterior and lateral margins, a yellow curved stripe, concave anterad, near base; elytra mostly yellow; aedeagus as in figures 21-23; length 3.5 - 4 N à a ee A cd se, PRI volkovitshi Pronotum black with anterior and lateral margins yellow; elytra red with dark pattern; length 4.0 HAFEN u u Sc ei A IT: ee RL N amasiensis 122 SASSI & KISMALI 42 Pronotum with 8 black spots (in two rows); elytra with 2 spots in posthumeral area and 2 spots on clivus; in the palest specs. the dark spots are mostly reduced; anal sternite in male with a pointed tubercle bordering a shallow depression; length 5.0 - 6.0 mm ........ quatuordecimmaculatus - Only 4 spots on pronotum; eytra with 2 spots in posthumeral area and 1 spot on clivus ... 43 43 Head completely black (sometimes labrum yellowish). Anal sternite in male with a little tran- sverse bifid ridge bordering the anterior margin of a shallow depression; length 5.0 - 5.5 mm . veci li acetone efile ehren Let ss ione dee gilteri - Head with a little yellow spot under eyes; anal sternite in male with a pointed tubercle; length Coira diporto) ps sata ten hrs epee els erage pseudoreitteri 444 Borsalesurtassentirchhionpatlyapetallion che ma encore a pa 45 Jee PD orsaleeumiceisepmictalllic he ber uses ee sie sonda 04 ar bn res 39 45. Only pronotum-metallic; elytra reddish with dark pattern ae, as, aaa mans ns 46 nel iaparbporeatitelyanzialbe ssnellnstetmen! na nun dona de cere x he 48 46 Only pronotum with long hairs; elytral surface bald; length 4.0 - 5.0 mm .......... pit lIChs -1Pionommandéltmlonehaismieue svasata. inabile peo pale 47 47 Punctures of pronotum strongly elongated, its surface appears rugose; elytra yellow with black spots, or almost entirely black in melanistic forms; length 4.0 - 5.0 mm .......... rugicollis 2 Puneturss:on pronotumroundedselytraweddish: mon deal a ara euchirus (female) 48 Dorsal surface with long hairs; abdominal sternites partly fused together; first abdominal ster- nite in male with a longitudinal process reaching anal sternite: subg. Protophysus (pars) .. 49 sensor Aetna sario botanici prio CR. arr ee anatase. Lo Guar a lay tages ds pil 49 Metatibiae in male specs. without apical enlargement; elytral surface in female mostly reddish; aedeagus and abdomen as in figures 24-27; length 5.5 - 6.2 mm (Iranian species, its presence in Turkeyäisipossible unsheseasternmestternitories).. sr a duo dea zur un euchirus - Metatibiae in male specs. with a leaf - shaped apical enlargement; elytral surface in female mostly meta MICAS re teed ts toi tral wel ted Belarus RS Qe dee à 50 50 Longitudinal process in 1st abdominal sternite in male not bifurcate on apex (figure 31); in fema- le reddish pattern limited at elytral apex; legs yellowish in female; aedeagus as in figure 28-30. Leone AN Fannie tendent brioche le ai i ARR OLA I à moehringi - Longitudinal process in first abdominal sternite of male bifurcate on apex (figure 35); in fema- le reddish pattern often reaches anterior half of elytra; at least femora entirely metallic in both sexes: Pedeaous as in.figtres 32-34, lenoth 3: 7-6, Zain ua. mana wes dialer owes el wehnkei 51 Larger species, length 6.0 mm or more, whole surface, except rare forms, green ......... 52 sor siunaller species) lensth less than 6.0: mam, surface ereenor blue... sus 1 il. 54 52 Male specs. with anal depression bounded by a bifid ridge. Pronotum usually stronger and den- ser punctured, surface between punctures more opaque; length6.0-7.5mm ....... sericeus 1 )Malespees: withoutmdees bordering anal. Cepressione: Lio Le arena it ou tal + 53 53 Margins of pronotum straighter and larger, usually completely visible when viewed from above; ligula of aedeagus long and pointed, exceeding fraena of endophallus; length 6.0 - 7.5 mm ... anid aust: hate eA breed cet een erie herd son acs verlassenen aureolus - Margins of pronotum more curved and slender, usually visible only near posterior angles when viewed from above; pronotal punctures finer and sparser, surface between punctures lustrous; ligula of aedeagus short, not exceeding fraena of endophallus; first sclerite of endophallus as in figures 39-42. Spermatheca as in figure 46; length 6.0 - 8.2 mm ............. paphlagonius 54 Dorsal surface deep blue to blackish, never shiny green; margins of pronotum more slender, impunc- tate; punctures of disc small and sparse (only male specs. identifiable for certain) .......... dI - Dorsal surface usually shiny green (rare forms metallic blue); margins of pronotum larger, punctu- red; disc of pronotum with coarse punctation (only male specs. identifiable for certain) ...... 58 The Cryptocephalinae of Turkey 123 55 56 87 64 65 66 Anal sternite in males with a shallow depression never bounded by any relief; surface of pro- notum more shiny, sometimes with green tints; apex of aedeagus (figure 51) shortly triangular; ventral surface deeply:sunken'toward'apexlength 4 3311 IF IE IR virens Anal sternite in male with a relief of various shape; apex of aedeagus long and narrow . . . .56 Anal sternite with a shallow depression whose anterior margin is bounded by a thin little ridge. Ventral surface of apex of aedeagus (figure 52) almost flat; length 5.0 - 6.0 mm (doubtful pre- sence di) Turkey) 4 er WA PALI RITARDI, AMAT + (RA at SR violaceus Anal depression in male specs. deep and bounded by a more prominent relief; ventral surface of apex of aedeagus with two longitudinal rounded carinae, bounding a slender central area; dor- suit with saoletté blackisiitinits UA REN Ae III LITRO IE 2 57 Anal depression with a v - shaped relief anterad; ventral surface of aedeagus (figure 49) split- ted from basal foramen by a distinct ridge; length 4.5 - 5.5 mm ................ duplicatus Anal depression with a pointed, bent tubercle anterad; ventral surface of aedeagus (figure 50) without posterior distinct ridge; length 4.5 - 5.4 mm (Iranian and Caucasian species, never recor- ded in Turkeyoaccordine- de Lopatn). <.< ran ne beeen ae ch pen in SE concolor Apex of aedeagus triangular, in lateral view not turned ventrally; anal depression in male dee- peas lengih4.0- 5 Ham EAN a NR oh SD HR ruguliventris Apex of aedeagus more elongated, in lateral view distinctly turned ventrally; anal depression in male shallower: length 4:05. Saat Fr ERTL FE A QU, ICH praticola Upper part covered with long hairs; pronotum black with metallic tints, its surface appears rugo- se; elytra yellow with dark spots, sometimes coalescent to form longitudinal stripes; in darkest form elytra completely black; length 4) ~$ mimi a DA IT SR IAA rugicollis Upper part bare 0: Ha MEN DDA N EPTO REP 60 Elytra black, yellow pattern, if present, only on apex and posthumeral margins .......... 61 Elytra mostly redor vello; Pr CORRE CIR REINA, BIER 64 Elytra completely blackrtenatir 4.0 = I 1 DAME signatifrons At least anterior part of lateral margins of ey a la Mate PO TR EE M 62 Apex of elytra yellow; head and legs completely black; length 4.5 - 5.0 mm . . quadriguttatus Apex of dytia black (perio A ii Salen ae aes 63 Pronotum more convex and less transverse, with anterior and lateral margins yellow; yellow lateral strip not reaching posterior angles; head generally entirely yellow; length 3.5 - 5.0 mm bay Be hah ER AMP N INI eat) ei Poe. Aura TL dl ina Pronotum less convex and more transverse, anterior and lateral margins yellow as well, but late- ral strips in male reaching posterior angles; head black with clipeus, genal spaces and a sub- triangular macula on frons yellow; length 3.0 - 4.0 mm ........................ turcicus Pronotum ochraceus, dull, deeply and coarsely punctured, generally with two little black flecks on disc; elytra yellow, strongly punctured, with four (2 + 2) or five (2 + 2 + 1) flecks (partly obsolete in palest forms) arranged in oblique lines; ventrites and legs yellow; length 4.5 - 5.5 WM ag sce ee NE eee ed aes Secs Sloe ees eee cee ee octomaculatus Not such a combination af characters. ua ae a er a a a EEE at 65 Lateral margins of pronotum very broad, forming a shallow gutter; elytral margin broad as well, entirely visible from above from humeri to clivus; pronotum black, anterior and lateral margins yellow; a yellow median stripe from anterior rim, sometimes forming an anchor-shaped pattern; elytra red with suture, lateral rims and three spots (2,1) on disc black; spots often more or less eoalescent or obsolete: . #08 weeded eis RR ORI EINER 66 Pronotal and elytral margins slimmer pic anal U IE 69 Elytral epipleura black, femora black with a whitish spot on apex ..................... 67 Elytral epipleura red, legs black, sometimes fore ublapalerı 1... VA FEAR ke & 68 124 SASSI & KISMALI 67 Body slender, humeral spot smaller, usually covering only 1/4 postmedian part of humeral tuber- cle; elytra generally with three spots (2 + 1); aedeagus as in figure 47; length 5.0 - 5.5 mm ... RTRT BAG ANS ohare BIN IRE NE Pa Syren’ Gran lei ss danse à el rar. sexpunctatus - Body stouter; humeral spot larger, usually extended on posterior half of humeral tubercle; ely- tra generally with 4 spots (2 + 2) sometimes coalescent; aedeagus as in figure 48; length 5.5 - CHORIN VOTO RTAS aes > I ASTE EOS Li Pee TINI octopunctatus 68 Prototum less transverse (length / width ratio about 0.65); elytral black pattern reduced; exter- nal anterior spot sometimes splitted or missing; aedeagus slimmer apically (figure 53); length CH) er Oro Sika sebbene bobenc» fail se asa aw rie biledjekensis - Pronotum more transverse (length / width ratio about 0.58); median yellow fimbria on prono- tum often anchor - shaped apically, in particular in female (= trapezensis form); black spots on elitra wider; aedeagus widened terminally; length 5.5 - 7.0 mm ................. cribratus 69 Slim, cylindrical; elytra red with four black spots (2,2) often coalescent and forming a postba- sal and a subapical transverse band, commissure black; aedeagus as in figure 55-57; length 6.0 EI DOT RL EEE POLE ENDE se ee ars erp ad sa | tappesi “Stoner elymaredavith:tworor three black spots; névericoalescent 1... vili i sue 70 70 Elytra usually with two (1 + 1) spots (sometimes a further spot in the anterior half); commissu- re black; pronotum black with lateral margins, a median longitudinal stripe and a heart-shaped spotuntenontomentellum yellow lea 3-6 I mint) a. NN RE in cordiger EB BREI EEE de tele. | 71 Male: pronotum black, lateral margins more slender and curved; protibiae curved, bisinuated on inner side; metafemora with large obtuse denticle on outer rim; in female anterior margin of pro- TOME eNO {Ae otis GON oO is pe ale ew TR UN LA a tn RS loebli - Male: pronotum black with yellow pattern, lateral margins larger; shape of legs unmodified; in roma anenommaeinis black OS PNR et, eas Pou wea de vey ca prusias ACKNOWLEDGEMENTS We are greatly indebted with Dr N. Berti (Muséum national d'Histoire naturelle, Paris), Dr K. Schneider (Martin-Luther-Universitat, Halle), Dr H. Schönmann (Naturhistorisches Museum, Wien), Dr H. Wendt (Museum fiir Naturkunde der Humboldt-Universität, Berlin), Dr L. Zerche (Deutsches Entomologisches Institut, Eberswalde) for allowing us to study types under their care. Additionally, we thank Dr I. Löbl (Muséum d’ Histoire naturelle, Geneva), Dr H. Silfverberg (Zoological Museum, University of Helsinki), Dr A. Vigna Taglianti (Universita “La Sapienza”, Rome), Dr J. Bezdek (Brno), Dr D. Erber (Giessen), Dr H. Kippenberg (Herzogenaurach), Dr M. Zuber (Kosmonosy), Mr S. Zoia (Milan) for loaning specimens. We are also grateful to Prof. I. Lopatin and Prof. A. Warchalowski who provided bibliographic data and unpublished informations that significantly improved the manu- script. We express our gratitude to Dr E. G. Riley (Texas A&M University) for the English langua- ge revision. Our cordial thanks are also due to Mr M. Pavesi (Museo di Storia Naturale, Milano) and Dr A. Gok (Suleyman Demirel University, Isparta) for the valuable remarks. REFERENCES APFELBECK V., 1901 - Bericht über eine entomologische Forschungsreise nach der Türkey und Griechenland im Jahre 1900. Wissenschaftliche Mitteilungen aus Bosnien und der Herzegowina, 8, 3: 145-167. ASLAN I. & OzBEK H., 1997a - New records of the subfamilies Cassidinae and Cryptocephalinae The Cryptocephalinae of Turkey 125 (Coleoptera, Chrysomelidae) for the Turkish fauna. Zoology in the Middle East, 14: 87-91. ASLAN I. & ÖZBEK H. 1997b - The Checklist of the subfamily Cryptocephalinae (Coleoptera, Chrysomelidae) in Turkey. Atatiirk Universitesi, Ziraat Fakiiltesi Dergisi, Erzurum, 28, 2: 235- 259. ASLAN I. & ÖZBEK H., 1998 - New records of the subfamily Cryptocephalinae (Coleoptera, Chrysomelidae) for the Turkish fauna. Zoology in the Middle East, 16: 81-84. BEDEL L., 1889-1901 - Faune des Coléoptères du bassin de la Seine. V. Phytophaga. Annales de la Societé Entomologique de France (hors série), 1-423. BODEMEYER B., 1900 - Quer durch Kleinasien in der Bulghar - Dagh. Eine wissenschaftliche Studienreise. Unter Mitarbeit von G. Ganglbauer, Karl Josef Daniel, J. Weise u. a. Emmendingen, 1-169. Berti N. & RAPILLY M., 1973 - Contribution a la faune de l'Iran. Voiages de MM. R. Naviaex et M. Rapilly (Col. Crysomelidae). Annales de la Societé Entomologique de France (N. S.), 9, 4: 861-894. BERTI N. & RAPILLY M., 1979 - Contribution a la faune de l'Iran. 4. Cryptocephalinae. Nouvelle Revue d’ Entomologie, IX, 3: 239-269. BREIT J., 1918 - Beitrag zur Kenntnis der Arten des Genus Cryptocephalus Geoffr. Wiener Entomologische Zeitung, 37, 1-3: 35-52. BURLINI M., 1956 - Revisione dei Cryptocephalus Italiani e della maggior parte delle specie di Europa (Col. Chrysomelidae). Memorie della Società Entomologica Italiana, 34 (1955): 5-287. BURLINI M., 1965 - II Contributo alla conoscenza dei Cryptocephalini paleartici. Cryptocephalus espanoli, n. sp., della Penisola Iberica. (Coleoptera Chrysomelidae). Bollettino della Società Entomologica Italiana, 95: 17-19. BURLINI M., 1968 - Revisione delle specie italiane e della maggior parte delle specie europee del genere Pachybrachis Chevr. (Coleoptera Chrysomelidae). XXII Contributo alla conoscenza dei Cryptocephalini. 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WEISE J., 1900a - Neue Coleopteren aus Kleinasien. Deutsche Entomologische Zeitschrift, 1: 132- 141. WEISE J., 1900b - Beschreibungen von Chrysomeliden und synonymische Bemerkungen. Archiv für Naturgeschichte, 1, 2: 267-294. WEISE J., 1901 - Nachtrag zum kleinasiatischer Coleopteren von Hrn E. v. Bodemeyer, Freiburg 1. Br. 1900. Deutsche Entomologische Zeitschrift, 2: 203-204. WEISE J., 1906a - Pachybrachis bodemeyeri. Deutsche Entomologische Zeitschrift, 2: 472. WEISE J., 1906b - In: BODEMEYER, E. v. 1906. Beiträge zur Käferfauna von Klein-Asien. Deutsche Entomologische Zeitschrift, 2: 417-434. WINKLER A., 1929 - Catalogus Coleopterorum regionis palaearcticae. Cryptocephalinae. Wien, 1249- 1265. Authors addresses: D. Sassi, via san Rocco 17, I-22030 Castelmarte CO, Italy. E-mail: d.sassı@pn.itnet.it S. Kismali, University of Ege, Faculty of Agriculture, Department of Plant Protection, Bornova, 35100 Izmir, Turkey. E-mail: kismali @ziraat.ege.edu.tr “il Te Hd: bstooliag’ (in hy ‘Off vi puerta di n dia; cu 2 i at A4 Bar: TÉ Li TRUE si . 7 à i t si a | 7 a: n lapidi ni a in Fouad py Vi ia Te ta ie SU ‘ hi ai A L 1 Mem. Soc. entom. ital., 78 (1): 131-166. 30 aprile 2000 Roberto CALDARA Revisione dei Pachytychius delle Regioni Afrotropicale e Orientale (Coleoptera Curculionidae) Riassunto - Viene effettuata la revisione delle specie afrotropicali e orientali del genere Pachytychius Jekel, 1861; si tratta di 15 specie, sette delle quali risultano nuove per la scienza: P. aethiopicus (Etiopia), P beardae (Zambia), P. cognatus (Africa nord-orientale, penisola arabica), P. lyali (Zambia), P. oberprieleri (Sud Africa), P. striatellus (Kenya) e P. tuberculatus (Mozambico, Sud Africa). Sono proposte le seguenti nuove sinonimie: P. congoanus Hustache, 1923 = P. elongatus (Gyllenhal, [1835]); P. setosus ssp. transmutatus Voss, 1962 = P. erythreensis Hustache, 1932. In base a nume- rosi caratteri morfologici, risulta verosimile che le specie prese in esame appartengano a tre grup- pi strettamente correlati fra loro filogeneticamente: il gruppo di P. aridicola (Wollaston) (com- prendente specie delle regioni paleartica meridionale e afrotropicale), il gruppo di P. haemato- cephalus (Gyllenhal) (comprendente specie paleartiche, afrotropicali e indiane) e il gruppo di P. albosparsus (Fahraeus)(comprendente solo due specie dell’ Africa meridionale). Abstract - Revision of Afrotropical and Oriental species of Pachytychius Jekel (Coleoptera Curculionidae) The Afrotropical and Oriental species of the genus Pachytychius Jekel, 1861 are treated systema- tically; 15 species are recognized, seven of them are new: P. aethiopicus (Ethiopia), P. beardae (Zambia), P. cognatus (North-eastern Africa, Arabian peninsula), P. lyali (Zambia), P. oberpriele- ri (South Africa), P. striatellus (Kenya), and P. tuberculatus (Mozambico, South Africa). The fol- lowing new synonymies are proposed: P. congoanus Hustache, 1923 = P. elongatus (Gyllenhal, [1835]); P. setosus ssp. transmutatus Voss, 1962 = P. erythreensis Hustache, 1932. Three phyloge- netically closely related groups are recognized: the P. aridicola group (including southern Palaearctic and Afrotropical species), the P. haematocephalus group (including Palaearctic, Afrotropical and Indian species), and the P. albosparsus group (including only two southern African species). A key to the species also in English is provided. Key words: Coleoptera, Curculionidae, Curculioninae, Pachytychius, Afrotropical Region, Oriental Region, taxonomical revision, new species. INTRODUZIONE Il genere Pachytychius Jekel, 1861 è largamente distribuito, con circa una quaranti- na di specie, nella Regione Paleartica, dalla penisola iberica all'Afghanistan (non si hanno dati sulla sua presenza in Cina e in Giappone). Esso è presente anche nell'Africa subsaha- riana e in India come testimoniato da poche descrizioni, per la precisione sette per la prima Regione e tre per la seconda. Mentre le specie della Regione Paleartica sono state sottoposte recentemente a revi- sione (Caldara, 1978), alla quale sono seguite piccole aggiunte (Caldara, 1984, 1988), quel- le delle altre Regioni non sono mai state prese in considerazione dopo la loro più o meno accurata descrizione. In questi ultimi anni ho svolto approfondite ricerche nei principali Musei europei allo 132 CALDARA scopo di rintracciare i tipi e di selezionare fra la miscellanea il maggior numero di esem- plari del genere Pachytychius raccolti nelle Regioni extra-paleartiche. A questi sono da aggiungere altri esemplari ricevuti in studio da numerosi Colleghi. Sono ora in grado di completare la revisione di questo genere a livello mondiale. MATERIALE E METODI Per la presente revisione ho esaminato materiale delle serie tipiche di tutte le specie precedentemente descritte delle Regioni Afrotropicale e Orientale. Come per le specie paleartiche (Caldara, 1978), ho riportato la descrizione di ogni taxon, effettuata su un esemplare rappresentante nei limiti del possibile la forma più comu- ne nell’ambito della variabilità, omettendo quelle caratteristiche comuni a tutto il genere. Inoltre, per mezzo di un micrometro oculare, ho effettuato le seguenti misurazioni, espres- se in valori assoluti o come rapporti: lunghezza totale in mm dall'apice del pronoto all'a- pice delle elitre lungo la linea mediana; lunghezza del rostro (Rlu) dal bordo anteriore del- l'occhio all'apice con esclusione delle mandibole; lunghezza del pronoto (Plu) dalla base all'apice lungo la linea mediana e sua larghezza (Pla) al punto più largo; lunghezza delle elitre (Elu) dalla base all'apice lungo la linea mediana, loro larghezza (Ela) al punto più largo. ACRONIMI ARCEH. Collezione privata Alexander Riedel, Friedberg, Germania. BMNH. Department of Entomology, The Natural History Museum, London, Inghilterra (C. Lyal, J. Beard). CWOB. Collezione privata Charles W. O'Brien, Entomology-Biological Control, Florida A. & M. University, Tallahassee, USA. DBAU. Dipartimento di Biologia Animale e dell'Uomo, Università degli Studi di Roma "La Sapienza", Roma, Italia (E. Colonnelli). DEIE. Deutsches Entomologisches Institut, Eberswalde, Germania (L. Behne). ISNB. Institut Royal des Sciences Naturelles, Bruxelles, Belgio (P. Dessart). HNHM. Hungarian Natural History Museum, Budapest, Ungheria (O. Merkl). MACT. Musee Royal de l'Afrique Centrale, Tervuren, Belgio (H. M. André). MHNG. Muséum d'Histoire Naturelle, Genève, Svizzera (C. Besuchet, J. Löbl). MNHB. Museum fiir Naturkunde der Humboldt-Universitat, Berlin, Germania (F. Hieke). MNHP. Muséum National d'Histoire Naturelle, Paris, Francia (H. Perrin). MSNM. Museo Civico di Storia Naturale, Milano, Italia (C. Leonardi, C. Pesarini). NHMB. Naturhistorisches Museum, Basel, Svizzera (M. Brancucci). NHRS. Naturhistoriska Riksmuseet, Stockholm, Svezia (B. Viklund). RCMC. Collezione privata Roberto Caldara, Milano, Italia. SAMC. South African Museum, Cape Town, Sud Africa (V. Whitehead). SANC. South African National Collections, Pretoria, Sud Africa (R. Oberprieler). USNM. National Museum of Natural History, Washington, USA (J. Pakaluk). ZMKB. Zoologische Forschunginstitut und Museum A. König, Bonn, Germania (M. Schmitt). Revisione dei Pachytychius afrotropicali e orientali 133 Pachytychius Jekel Pachytychius Jekel, 1861: 270 (specie tipo: Tychius sparsutus (Olivier, 1807), per designa- zione originale). Tournier, 1874: 452; 1874a: 86. Gonzalez, 1968: 107. Caldara, 1978: 133. Styphlotychius Jekel, 1861: 271 (specie tipo: Tychius scabricollis Rosenhauer, 1856, per desi- gnazione originale). Tournier, 1874: 456; 1874a: 87. Caldara, 1978: 133. Barytychius Jekel, 1861: 272 (specie tipo: Tychius hordei Brullè, 1832, per designazione ori- ginale). Tournier, 1874: 460; 1874a: 69, 90. Caldara, 1978: 133. Rabdotorhinus Desbrochers, 1894: 82 (specie tipo: Rabdotorhinus hircus Desbrochers, 1894, per monotipia). Caldara, 1978: 133. DESCRIZIONE DEL GENERE. Tegumenti da neri a rossastri, ricoperti da squame variabili per forma e numero. Rostro subcilindrico più o meno arcuato, a scrobe che nascono oltre la metà e si dirigono in basso obliquamente all'asse del rostro, visibili di lato solo nella parte anteriore. Antenne rossastre; apice dello scapo, in posizione di riposo, raggiungente il mar- gine anteriore dell'occhio; funicolo di 7 segmenti, il primo allungato, il secondo sempre | più lungo che largo, ma più corto e più esile del primo, gli altri via via piu corti fino a distin- tamente trasversi. Occhi non debordanti dalla convessità del capo. Protorace più o meno trasverso, con lobi oculari ben sviluppati. Prosterno con margine anteriore più o meno inca- vato, ma sempre senza solco mediano. Scutello da invisibile a ben sviluppato. Elitre a lati da paralleli ad arrotondati, da globose ad allungate. Zampe con femori clavati, i posterio- ri a volte con un dente; tibie con margine interno senza dentini; tarsi con terzo segmento, di solito, distintamente bilobato; unghie semplici. Il 3 si distingue dalla 2 essenzialmente per il rostro lievemente più corto, ma soprat- tutto più fortemente carenato e punteggiato, per l'addome più appiattito e con più eviden- te concavità mediana lungo gli sterniti I e II. POSIZIONE SISTEMATICA DEL GENERE. Fino a pochi anni fa il genere Pachytychius è sempre stato posto nella sottofamiglia Erirhininae, tribù Erirhinini. Tuttavia, recentemente tale sot- tofamiglia è stata oggetto di approfondita revisione (Thompson, 1992; Kuschel, 1995) e, sulla base della morfologia degli organi genitali maschili, è stata scissa in raggruppamen- ti molto distinti dal punto di vista filogenetico. Tutti i generi caratterizzati da lobo media- no di tipo "ortocero", cioè con lamina dorsale tegminale ben sviluppata (ad esempio come in Attelabidae e Apionidae), carattere ritenuto plesiomorfo, sono rimasti nella sottofami- glia Erirhininae (Kuschel, 1995) o formano ora la famiglia Erirhinidae (Thompson, 1992), mentre gli altri generi caratterizzati da parameri del tegmen ridotti, carattere considerato apomorfo, sono stati collocati in varie tribù della sottofamiglia Curculioninae. Per quanto riguarda Pachytychius non è ancora chiara la tribù di appartenenza: Alonso-Zarazaga (comun. pers.) ipotizza l'inclusione del genere nella tribù Storeini. DIVISIONE DELLE SPECIE IN GRUPPI. Nella revisione dei Pachytychius della Regione Paleartica (Caldara, 1978), avevo effettuato un tentativo di divisione del genere in gruppi di specie sulla base sia di caratteristiche morfologiche esterne che del lobo mediano dell’edeago. In seguito, ho cercato di sviluppare un approccio di ricostruzione filogenetica del genere su base cladistica usando come sister-group il genere Aubeonymus Jacquelin du Val, 1854. I dati preliminari desunti da tale studio, tuttora in corso, sembrano avvalorare nel comples- so i gruppi come da me inizialmente intesi. 134 CALDARA Nelle regioni prese in esame dal presente studio, le specie appartengono a soli tre gruppi, due dei quali già delineati in base allo studio delle specie paleartiche ed uno apparente- mente endemico dell’ Africa meridionale, strettamente imparentati fra loro e facenti vero- similmente parte di un'unica linea evolutiva, come sembrerebbero dimostrare le marcate similarità nella forma degli organi genitali. TABELLA DICOTOMICA DELLE SPECIE la 2 > 10. SPE Cie della Re sion. Alrotrapieale on. a. brani chie) aio nen gare ar 2 Speciali MONS Oil cut ei RT ni dut ral ar 15 Elitre allungate, da subrettangolari a debolmente subellittiche (figg. 1-4) ................. 3 Blitze cence subovalin gp pordietalbo span us ne IR Aa 12 Elitre con squame di due colori solitamente ben contrastanti fra loro, in ogni caso le più scure ricoprenti gran parte delle prime due interstrie da dove si dipartono a formare fasce e chiazze più o meno numerose e più o meno lunghe (gruppo di P. haematocephalus, figg. 3-4) .......... 4 Elitre con squame unicolori o di due colori poco contrastanti fra loro, in quest'ultimo caso le squa- me più chiare formano delle strisce più o meno complete su alcune interstrie dispari (gruppo di ERI 2 Oia EIN RUE 16000) tai] 8 Pronoto con il punto pit largo alla meta da dove si restringe lievemente e gradualmente, in modo subrettilineo, fino in prossimità dell'apice e bruscamente solo all'apice (fig. 32) ............. RAS in Ir ini ie dg 6. P. elongatus (Gyllenhal) Pronoto con il punto più largo alla metà o oltre da dove si restringe bruscamente fino all’ apice RE IL PE ATO TEE EAT KEIN, BERN EL A De IO FAR SÌ Pronoto distintamente convesso sul disco e a lati fortemente arrotondati (fig. 34). Interstrie eli- trali, soprattutto nel terzo apicale, con una Serie mediana irregolare di squame lievemente solle- vate dai tesumenti, visibili guardando Te elitre di profilo Ml: 8. P. lyali n. sp. Pronoto moderatamente convesso sul disco e a lati moderatamente arrotondati (figg. 31, 33). Interstrie elitrali tutt'al più con isolate squame lievemente sollevate dai tegumenti .......... 6 Disco del pronoto e prime due interstrie elitrali ricoperte da squame distintamente più scure delle altre. Tegumenti di protorace e di gran parte dell'addome nerastri. Pronoto fortemente e brusca- On Er SION on lege li carol a wy ES e ED CE EC RE NE 7 Disco del pronoto e prime due interstrie elitrali ricoperti da squame circa dello stesso colore delle altre. Tegumenti di protorace e addome completamente bruno-rossastri. Pronoto gradualmente MMe Ome enzo CA le (08x39 eo one ata cent 9. P. beardae n. sp. Elitre più lunghe (Elu/Ela 1,54-1,58). Pronoto con il punto più largo nel terzo anteriore ...... We Bo ee a ere te ee ee 7. P. leucoloma Jekel Elitre più corte (Elu/Ela 1,45-1,48). Pronoto con il punto più largo alla metà ............... ne Me to A nahi RUE RU de ee MAR ER ARE RUE RS IDRA RAS LICEI gilde 10. P. oberprieleri n. sp. Pronoto e soprattutto elitre con numerose squame strette o setole distintamente sollevate (figg. 172): Femor POserion Neo CON PICCOlO GENIE RE ee ea 9 Pronoto ed elitre con alcune squame tutt'al più debolmente sollevate. Femori posteriori con robu- ROIO ENO ROTTE NE 4 ry TTT oe vos 11 Femori posteriori inermi. Rivestimento dorsale costituito da squame setoliformi da moderata- mente a molto allungate, in parte subcoricate (lu/la 5-7) e in parte sollevate (lu/la 8-9); queste ultime sulle elitre sono inclinate di 30-45 gradi rispetto al piano dorsale e sono ricurve all'indie- libra a PU Na e lle live ar 3 1. P cognatus n. sp. Femori posteriori con piccolo dente. Rivestimento dorsale costituito da squame setoliformi più allungate, in parte subcoricate (lu/la 7-10) e in parte sollevate (lu/la 10-15); queste ultime sulle elitre sono inclinate di 60-90 gradi rispetto al piano dorsale e sono diritte ................ 10 Rivestimento elitrale formato in massima parte da squame biancastre; ogni interstria presenta 4- 5 serie Irresolari di aguamie subeoncate (ie. PD) i... de nat à La à 2. P. setosus Marshall Rivestimento elitrale formato da squame brune e squame biancastre; ogni interstria presenta 6- Revisione dei Pachytychius afrotropicali e orientali 135 id. 12: 13. 14. 7. sere irregolari di Squamiesobeonieae di Dane ZIEL ZA cee 3. P. erythreensis Hustache Pronoto con tre strisce strette di squame biancastre ben evidenti e ben delimitate, una centrale e due fra centro e lati. Rivestimento elitrale vittato. Pronoto moderatamente trasverso (Pla/Plu 1:2) rione br ae Is ie nie. ee patio arie np 5. P. striatellus n. sp. Rivestimento dorsale pressoché uniforme bianco-grigiastro. Pronoto fortemente trasverso (Pla/Plu WAS) osha duo dense A ee ial eleva] Lol te: as oe gl A 4. P. aethiopicus n. sp. Elitre moderatamente convesse, con quarta, quinta, sesta e settima interstria elitrale fortemente convesse e formanti una protuberanza ottusa a livello della declivita posteriore in vicinanza del loro apice, con omeri debolmente prominenti in avanti. Pronoto distintamente convesso, a lati NOM Carenaty ITS, D). ci idonea sake Ws ae a ee ae 15. P. tuberculatus n. sp. Elitre fortemente convesse, con quarta, quinta, sesta e settima interstria elitrale di uguale con- vessita fino all'apice. con omeri fortemente prominenti in avanti. Pronoto appiattito soprattutto verso ilativiche sono Care HEN IE URRA ee a 14. P. albosparsus (Fahraeus) Dimensioni minori (lunghezza mm 2,6-3,2). Pronoto debolmente trasverso (Pla/Plu 1,15-1,20). Femori posteriori con piccolo dente. Sulle elitre le squame più scure ricoprono tutt’al più le prime due interstrie e formano qualche confusa chiazza sulle altre interstrie (fig. 4) ............... inn no ir anioni ee i enon vie 13. P. viciae Marshall Dimensioni maggiori (lunghezza mm 3,9-5,0). Pronoto da moderatamente a fortemente trasver- so (Pla/Plu 1,30-1,45). Femori posteriori con grosso dente. Sulle elitre le squame più scure rico- prono le prime due interstrie e formano numerose chiazze sulle altre interstrie............ 14 Pronoto meno trasverso (Pla/Plu 1,30-1,35), ricoperto quasi interamente da squame scure, alcu- ne più chiare formano una confusa e stretta striscia longitudinale mediana e piccolissime chiaz- zeite verso lati ul is tee ete pepe > bestiali ventre rh rene 12. P mungonis Marshall Pronoto più trasverso (Pla/Plu 1,40-1,45), ricoperto almeno per la metà da squame chiare sia sul disco che al lati „Bass; ei a CERN UM ETAGE RE 11. P indicus Tournier KEY TO THE SPECIES bn Del Species from the Afötropie REIST TEE RA HELEN Ee eed A ee nae, 2 Species fromthe Oriental Region Teen A a RIT IRA aa 13 Elytra elongate, subrectangular to weakly subelliptical (figs. 1-4) ....................... 3 Elytra short suboval (PF albosparsus’ Mon ter S=G) aie N a On à. 12 Elytra covered with scales which are usually of two distinctly different colours, with the darker covering most part of interstriae 1 and 2 and forming more-or-less numerous and more-or-less elongate transverse bands and spots (P. haematocephalus group, figs. 3-4) ................ + Elytra covered with scales which are unicolorous or of two scarcely different colours, in this lat- ter case with the lighter scales forming more-or-less complete vittae on part of odd-numbered interstriae (P. aridicola group, fes. 132) ke ali i be an ne 8 Pronotum widest at middle, gradually and rectilinearly narrowing to apex proximity and stron- gly narrowing only at apex IE. 2 LS er 6. P. elongatus (Gyllenhal) Pronotum widest at middle or at after middle, strongly narrowing to apex (figs. 31, 33-34) ...5 Pronotum distinctly convex on disc and with sides distinctly rounded (fig. 34); elytral interstriae, especially in apical third, with irregular median row of scales slightly raised and visible from lateral VAS no: xo so rr niece eases I I E n 148. P. tyali n. SD. Pronotum moderately convex on disc and with sides moderately rounded (figs. 31, 33); elytral interstriae at most with sparse Seales slightly ICP NIE EN ERBEN IF 6 Disc of pronotum and elytral interstriae | and 2 covered with scales distinctly darker than others; integument of prothorax and of most part of abdomen blackish; pronotum strongly and abruptly narrowed.at apex (ia AlN 5 iii i e Dare a eae 7 Disc of pronotum and elytral interstriae 1 and 2 covered with scales only slightly darker than others; integument of prothorax and abdomen completely brown reddish; pronotum gradually narrowing in apical third (fie. 33) 124 IN ae ME ER DIRE 9. P. beardae n. sp. Elytra longer (length/width 1,54-1,58); pronotum widest at apical third . .7. P. leucoloma Jekel 136 CALDARA - Elytra shorter (length/width 1,45-1,48); pronotum widest at middle . . .10. P. oberprieleri n. sp. 8. Pronotum and especially elytra with numerous narrow scales or setae distinctly raised (figs. 1- lhındıie moranunanınedrerwithksmalltooth HN. 22, DTA AE DI 9 - Pronotum and elytra with some scales at most slightly raised; hind femora with robust tooth 11 9. Hind femora unarmed; scales forming dorsal vestiture moderately to distinctly elongate (subre- cumbent scales length/width 5-7, raised scales length/width 8-9); raised elytral scales at 30-45 destees im telation to dorsal plane and backward! mn. eroi 1. P. cognatus n. sp. - Hind femora with small tooth; scales forming dorsal vestiture more elongate (subrecumbent sca- les length/width 7-10, raised scales length/width 10-15); raised elytral scales at 60-90 degrees in fl AMOMMOMOnsAl lane and serata 2 RT er II 10 10. Elytral vestiture mostly composed of whitish scales; subrecumbent scales less adpressed (4-5 ie sularseniesiloreachanterst Ha aie) Is IE Se ad OUR 2. P. setosus Marshall - Elytral vestiture composed of brown and whitish scales; subrecumbent scales more adpressed (6- Aine cular series fOmeacinmtensunmayiie 2). NR TO UE 3. P. erythreensis Hustache 11. Pronotum with three distint vittae of white scales, one midline and two laterally; elytral vestitu- re vittate; pronotum moderately transverse (width/length 1,21) ......... 5. P. striatellus n. sp. - Dorsal vestiture nearly unicolorous, greyish-white; pronotum strongly transverse (width/length1,45) Re NT RUNTER SN IRAP DIMARO A CARS ES. SUR mE REED Ui DAS DROIT LI LIO, 4. P. aethiopicus n. sp. 12. Elytra moderately convex, with interstriae 4, 5, 6 and 7 at apex proximity markedly convex and forming blunt tubercle, with humeri weakly prominent anteriorly; pronotum distinctly convex, NOLA alate MONETARIE DRITTEN EN PIE 15. P. tuberculatus n. sp. - Elytra strongly convex, with interstriae 4, 5, 6 and 7 of same convexity to apex, with humeri strongly prominent anteriorly; pronotum flattened especially toward sides, which are carinate (iero scali) Pores ssa cioniti Saga sbk be ote 14. P. albosparsus (Fahraeus) 13. Body smaller (length mm 2.6-3.2); pronotum weakly transverse (width/length 1,15-1,20); hind femora with small tooth; on elytra dark scales covering at most interstriae 1 and 2 and forming Some COMMSEd SPOls'ON OlMEIS (setti e ea e Eee da 13. P. viciae Marshall - Body larger (length mm 3.9-5.0); pronotum moderately to strongly transverse (width/length 1,30- 1,45); hind femora with robust tooth; on elytra dark scales covering interstriae 1 and 2 and for- TEA INC TOUS POLONIA AES ER I e RAIL Ing dora en et à 14 14. Pronotum less transverse (width/length 1,30-1,35), covered nearly completely with dark scales, with some lighter scales forming a confused and narrow median longitudinal vitta and very small SOC ORTE AA INCI SAGRA] rae agi Jive ne ER Ka e RE TR 12. P. mungonis Marshall - Pronotum more transverse (width/length 1,40-1,45), at least half of which covered with light sca- Nie SD ONO TA Tela Os ATC GIST 1 tage a EE aci 11. P indicus Tournier ELENCO DELLE SPECIE Gruppo di Pachytychius aridicola Il gruppo presenta la combinazione dei seguenti caratteri: elitre allungate, da subret- tangolari a debolmente subellittiche, ricoperte da squame solitamente allungate, da subret- tangolari a decisamente piliformi, spesso in parte distintamente sollevate dai tegumenti, di colore chiaro e disposte uniformemente o formanti strisce sulle interstrie dispari; femori da inermi a fortemente dentati; lobo mediano a forma di corto tubo. A questo gruppo appartengono tre specie della Regione Paleartica meridionale e cin- que specie della Regione Afrotropicale (distribuzione in fig. 35). Il gruppo appare stretta- mente correlato filogeneticamente con quello di P. haematocephalus, dal quale differisce essenzialmente per la diversa forma e disposizione delle squame del rivestimento elitrale (vedi tabella dicotomica). Revisione dei Pachytychius afrotropicali e orientali 157 1. Pachytychius cognatus n. sp. (figg. 7, 20, 25) DIAGNOSI. Un Pachytychius del gruppo aridicola con interstrie elitrali ricoperte da squa- me prevalentemente biancastre, moderatamente allungate, disposte in 4-5 serie irregolari, in parte subcoricate e in parte sollevate di 30-45 gradi rispetto al piano dorsale e ricurve all’indietro, con pronoto moderatamente trasverso, con femori posteriori inermi. LOCALITÀ TIPICA. Gibuti, El Hadj. SERIE TIPICA. Holotypus d: [Gibuti] “El Hadj, Dr. Martin / Sig. R. Oberthür (Coll. C. Martin), Eing. Nr. 4, 1956” (ZMKB).Paratypi: 1 ¢ “Gizan am Roten Meer, 25-26.III.1983 / Saudi Arabia, C. Holzschuh” (NHMB); 1 3 “Yemen, 1 mile W. of Ta’izz, on road to Mocha, ca. 4,500 ft., 20.x11.1937 / From Indigofera bushes / B.M. Exp. to S.W. Arabia, H. Scott & E.B. Britton, B.M. 1938-246” (BMNH); 1 6 “W. Aden Prot., Wadi Tiban, N.W. of Jebel Jihaf, ca. 3,800 ft., 22.x.1937 / Beaten from Indigofera bushes / B.M. Exp. to S.W. Arabia, H. Scott & E.B. Britton, B.M. 1938-246 / Pachytychius, J Balfour-Browne” (BMNH); 1 2 “Sudan, Wad Medani a Bl. Nil, 22.10.1979, lux, leg. Hieke” (MNHB); 1 9, idem eccetto “23.10.1979” (RCCM). DESCRIZIONE. 3 (holotypus). Lunghezza mm 2,7. Tegumenti interamente bruni; sul dorso ben visibili fra il rivestimento formato da squame moderatamente allungate (lu/la 5-7), strettamente lanceolate, biancastre, da cori- cate a moderatamente sollevate, disposte in 4-5 serie molto irregolari per ogni interstria; squame un poco più lunghe (lu/la 8-9) e più sollevate formano una serie abbastanza rego- lare su ogni interstria elitrale. Strie elitrali con una serie poco evidente di squame più pic- cole e strette di quelle delle interstrie. Parte ventrale ricoperta abbastanza fittamente da squame biancastre lunghe (lu/la 7-9), da coricate a subcoricate. Rostro un poco più lungo del pronoto (Rlu/Plu 1,16), visto dorsalmente a lati paral- leli, visto di lato distintamente e regolarmente arcuato, debolmente ristretto dall'inserzio- ne delle antenne all'apice. Antenne inserite fra terzo medio e terzo apicale; primo segmen- to del funicolo più robusto e circa 1,3 volte più lungo del secondo. Fronte un poco più stret- ta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,28), a lati moderatamente arrotondati, con il punto di maggior larghezza alla metà, moderatamente convesso, con punteggiatura fitta e abbastanza regolare, con intervalli fra i punti stretti e lucidi. Lobi oculari moderata- mente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben visibile. Elitre allungate (Elu/Ela 1,52), a lati subparalleli, con margine anteriore moderata- mente concavo, con omeri non prominenti in avanti e subarrotondati, discretamente più larghe del pronoto (Ela/Pla 1,32), moderatamente convesse. Interstrie subpiane; strie for- mate da punti grossi e profondi, debolmente visibili fra il rivestimento e larghe poco meno delle interstrie. Femori posteriori inermi; terzo segmento tarsale bilobato e moderatamente più largo del secondo. Edeago come in fig. 7. ® (paratypus). Rostro un poco più lungo (Rlu/Plu 1,24). Spiculum ventrale e sper- mateca come in figg. 20 e 25. VARIABILITÀ. Per quanto riguarda i paratypi, le dimensioni variano da 2,6 a 3,0 mm. Sul 138 CALDARA pronoto può essere più o meno evidente una stretta striscia longitudinale mediana di squa- me un poco più bianca delle altre, mentre squame brunastre possono essere presenti verso i lati o alla base in regione paramediana, dove formano una piccola chiazza poco eviden- te. Anche sulle elitre squame brunastre possono formare chiazzette più o meno numerose e più o meno evidenti, mentre le squame biancastre possono essere più fitte alla base di terza e quinta interstria e in regione omerale. Rlu/Plu d 1,10-1,18, 9 1,22-1,27; Pla/Plu 1,25-1,32; Elu/Ela 1,49-1,56; Ela/Pla 1,29-1,35. ETIMOLOGIA. L'aggettivo latino "cognatus" (vicino, imparentato) vuole evidenziare la stret- ta parentela della specie con P. setosus. CONSIDERAZIONI E NOTE COMPARATIVE. Specie strettamente imparentata con P. setosus, e con tale nome classificata da Voss (1962), ma sicuramente distinta da questa per la costan- te differenza nel tipo di rivestimento dorsale (vedi tabella). Per il rivestimento del prono- to, a volte caratterizzato da due chiazze più scure poco evidenti alla base, si può avvicina- re a P. basimaculatus Voss, 1964, anch'esso diffuso nel Sudan (Caldara, 1978), dal quale differisce per le minori dimensioni (mm 2,6-3,0 invece di mm 3,7-4,2), per le elitre distin- tamente più corte e per la serie di squame un poco sollevate sulla parte mediana delle inter- strie elitrali. NOTE BIOLOGICHE. Come riportato sui cartellini (vedi sopra), due paratypi risultano raccolti battendo rami di Indigofera sp. (Fabaceae) in due località differenti. DISTRIBUZIONE. Sudan, Gibuti, Yemen, Arabia Saudita. MATERIALE ESAMINATO. Oltre agli esemplari della serie tipica (vedi sopra), 1 d etichettato “Arabia, Buraim. 19-20.v.1936, H.St.J.B. Philby, B.M. 1936-521” (BMNH), che non ho considerato come paratypus perché un poco differente dagli altri esemplari a causa dei lati di pronoto ed elitre distintamente più arrotondati. 2. Pachytychius setosus Marshall (figg. 1, 8) Pachytychius setosus Marshall, 1950: 209. DIAGNOSI. Un Pachytychius del gruppo aridicola con interstrie elitrali ricoperte da squa- me prevalentemente biancastre, molto lunghe, disposte in 4-5 serie irregolari, in parte sub- coricate e in parte sollevate di 60-90 gradi rispetto al piano dorsale e diritte, con pronoto moderatamente trasverso, con femori posteriori con piccolo dente. LOCALITÀ TIPICA. Niger, Aîr, Agadez. SERIE TIPICA. Specie descritta su due esemplari del Niger (Air: Agadez e Monti Beguezans). Non ho esaminato l'holotypus di Agadez conservato nell'Institut Français d'Afrique Noire di Dakar, mentre ho potuto studiare il paratypus etichettato "Cotype / Irabellaben, Mts. Baguezans, 1200-1300 m, 26- 31.VIII.1947, Ifan, L. Chopard A. Villiers / Pachytychius setosus Mshl., Cotype £" (BMNH). RIDESCRIZIONE. d. Lunghezza mm 3,3. Tegumenti interamente bruno-rossastri; sul dorso ben visibili fra il rivestimento for- mato da squame molto lunghe (lu/la 7-9), strettamente lanceolate, biancastre e in minima parte bruno-grigiastre, da coricate a moderatamente sollevate, ricoprenti il pronoto (dove le biancastre formano tre fasce longitudinali moderatamente evidenti, una centrale più stret- Revisione dei Pachytychius afrotropicali e orientali 159 ta e due laterali) e le interstrie elitrali (dove sono disposte irregolarmente, grosso modo in 4-5 serie, e dove le biancastre ricoprono prevalentemente le interstrie dispari dando all'in- setto un aspetto vagamente vittato), e distintamente allungate (lu/la 10-12), piliformi, sube- rette, formanti una serie centrale irregolare su ogni interstria elitrale. Strie con una serie di squame molto sottili, ma più corte di quelle delle interstrie. Parte ventrale ricoperta abba- stanza fittamente da squame biancastre, lunghe (lu/la 9-12), da coricate a suberette. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,15), visto dorsalmente a lati subparalleli, debolmente divergenti all'apice, visto di lato distintamente e regolarmente arcuato, debolmente ristretto nella parte apicale. Antenne inserite fra terzo medio e terzo apicale; primo segmento del funicolo un poco più robusto e circa 1,2 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,34), a lati moderatamente arrotondati, con il punto di maggior larghezza alla metà, moderatamente convesso, con punteggiatura fitta e abbastanza regolare, con intervalli fra i punti stretti e moderatamente lucidi. Lobi oculari moderatamente pronunciati. Prosterno debolmente incavato nel mezzo. Scutello grande, ben visibile. Elitre allungate (Elu/Ela 1,52), a lati molto debolmente curvilinei nei due terzi ante- riori, con margine anteriore moderatamente concavo, con omeri non prominenti in avanti e subarrotondati, discretamente più larghe del pronoto (Ela/Pla 1,33), debolmente conves- se. Interstrie subpiane; strie formate da punti grossi e profondi, debolmente visibili fra il rivestimento e larghe poco meno delle interstrie. Femori posteriori con un piccolo dente aguzzo; terzo segmento tarsale bilobato e moderatamente più largo del secondo. Edeago come in fig. 8. 2. Rostro un poco più lungo (Rlu/Plu 1,22). Spiculum ventrale e spermateca come in P. cognatus (figg. 20 e 25). VARIABILITÀ. La lunghezza è compresa fra 2,7 e 3,7 mm. Il pronoto varia discretamente per larghezza. Rlu/Plu 4 1,12-1,16, 2 1,21-1,26; Pla/Plu 1,27-1,36; Elu/Ela 1,50-1,58; Ela/Pla 1,268 1535, CONSIDERAZIONI E NOTE COMPARATIVE. Specie intermedia fra P. cognatus e P. erythreensis, dai quali si separa essenzialmente per la differente disposizione del rivestimento elitrale (vedi tabella), caratterizzato da una serie di squame setoliformi più sollevate e diritte che in P. cognatus, ma meno erette e meno piliformi che in P. erythreensis. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Niger, Chad, Sudan, Etiopia, Yemen, Arabia Saudita. MATERIALE ESAMINATO. Oltre al paratypus (vedi sopra), altri 46 esemplari. Chad: Tchad, Distr. Kanem, N' Gouri, X/XI.1958, P. Renaud, ex coll. Dr. Breuning (3, MACT). Sudan: Sudan, Prov. N. Darfur, El Fasher (3, MNHB; 2, HNHM); Brit. Sudan, Khartoum, 15.X.1924, N. B. Johnson (1, BMNH); Sudan Govt., Khartoum, H. W. Bedford, 9.IX.1928, At light (1, BMNH); Sudan, Kutum, m 1140 (3, HNHM); Sudan, Senaar a Bl. Nil, lux (3, MNHB); Sudan, Wad Medani a BI. Nil, 29.X.1979, lux, leg. Hieke (4, BMNH; 20, MNHB; 3, RCCM). Etiopia: Etiopia, Bati (1, HNHM). Yemen: Arabia merid., Yemen, leg. Scortecci, Est di Rada, Rocce Nere, m 1880, 22.IX.1965 (1, MSNM). Arabia Saudita: Sahl Rakbah, 2-3.VI.1982, Saudi Arabien, W. Buttiker (1, NHMB). 140 CALDARA 3. Pachytychius erythreensis Hustache (figg. 2, 9) Pachytychius erythreensis Hustache, 1932: 43. Pachytychius setosus subsp. transmutatus Voss, 1962: 197 n. syn. DIAGNOSI. Un Pachytychius del gruppo aridicola con interstrie elitrali ricoperte da squa- me brune e biancastre, fortemente allungate, piliformi, disposte in 6-7 serie irregolari, in parte subcoricate e in parte sollevate di 60-90 gradi rispetto al piano dorsale e diritte, con pronoto moderatamente trasverso, con femori posteriori con piccolo dente. LOCALITÀ TIPICA. Fritrea, Adua. SERIE TIPICA. Specie descritta su tre esemplari raccolti ad Adua in Eritrea (Hustache non designa l’ho- lotypus) e dei quali ne ho esaminati due (4 ¢, DEIE), uno etichettato “Coll. Pape / Adua, Eritrea / Pachytychius erythreensis m. type, Hustache det. / Holotypus” (lectotypus qui designato) e l’altro “Coll. Pape / Adua, Eritrea / Hustache det. / Paratypus” (paralectotypus). Ä SINONIMI. Voss descrive la subsp. transmutatus di P. setosus su un unico esemplare raccolto in Gibuti paragonandolo a P. erythreensis. Ho esaminato tale esemplare (ZMKB) etichet- tato "La Sarrat, Dr. Martin / Chemin de fer Djibouti-Harrar Kilom. 163 / Holotypus" che non mostra nessuna sostanziale differenza dagli esemplari della serie tipica di P. eryth- reensis. RIDESCRIZIONE. d. Lunghezza mm 3,0. Tegumenti bruno-rossastri ad eccezione del protorace bruno-nerastro; sul dorso un poco visibili fra 1l rivestimento formato da squame molto lunghe (lu/la 7-10), strettamen- te lanceolate, biancastre e brune, da coricate a lievemente sollevate, ricoprenti il pronoto (dove le biancastre formano una stretta fascia longitudinale mediana) e le interstrie elitra- li (dove sono fitte, disposte irregolarmente e in parte un poco embricate, e dove le bianca- stre ricoprono la base della terza interstria, gli omeri e formano alcune chiazzette sparse poco evidenti), e fortemente allungate (lu/la 13-15), piliformi, da suberette a erette, for- manti una serie centrale abbastanza regolare su ogni interstria elitrale. Strie con una serie di squame molto sottili, ma più corte di quelle delle interstrie. Parte ventrale ricoperta abba- stanza fittamente da squame brune lunghe (lu/la 7-9), da coricate a subcoricate. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,17), visto dorsalmente a lati subparalleli, visto di lato distintamente ricurvo nel terzo basale, moderatamente arcuato nei due terzi apicali, debolmente e gradualmente ristretto dalla base all'apice. Antenne inseri- te fra terzo medio e terzo apicale; primo segmento del funicolo un poco più robusto e circa 1,5 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,20), a lati un poco arrotondati dalla base fino in prossimità dell'apice dove si restringono bruscamente e angolosamente, con il punto di maggior larghezza fra terzo medio e terzo apicale, discretamente convesso, con punteg- giatura fitta e abbastanza regolare, con intervalli fra i punti stretti e lucidi. Lobi oculari moderatamente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben visibile. Elitre allungate (Elu/Ela 1,45), a lati molto debolmente curvilinei nei due terzi ante- riori (Ela/Pla 1,35), con margine anteriore moderatamente concavo, con omeri non promi- nenti in avanti e subarrotondati, discretamente più larghe del pronoto (Ela/Pla 1,36), mode- ratamente convesse. Interstrie subpiane; strie formate da punti grossi e profondi, debol- Revisione dei Pachytychius afrotropicali e orientali 141 4 CL 5 etna gite La in Figg. 1-6. Habitus di: 1 — Pachytychius setosus Marshall; 2 — P. erythreensis Hustache; 3 — P. leuco- loma Jekel; 4 — P. viciae Marshall; 5 — P. albosparsus (Fahraeus); 6 — P. tuberculatus n. sp. Non in scala. mente visibili fra il rivestimento e larghe poco meno delle interstrie. Femori posteriori con un piccolo dente aguzzo; terzo segmento tarsale bilobato e moderatamente più largo del secondo. Edeago come in fig. 9. 2. Rostro un poco più lungo (Rlu/Plu 1,21). Spiculum ventrale e spermateca come in P. cognatus (figg. 20 e 25). VARIABILITÀ. La lunghezza è compresa fra 2,6 e 3,5 mm (il lectotypus misura mm 3,4). A parte le dimensioni nessun'altra variazione degna di nota negli esemplari esaminati. Rlu/Plu 142 CALDARA 3 1,15-1,18, 2 1,20-1,25; Pla/Plu 1,19-1,25; Elu/Ela 1,44-1,47; Ela/Pla 1,33-1,38. CONSIDERAZIONI E NOTE COMPARATIVE. Per la presenza di una serie di lunghe setole pilifor- mi erette sulle interstrie elitrali, la specie può essere confusa esclusivamente con P. seto- sus, le cui squame setoliformi sollevate dai tegumenti elitrali sono tuttavia in massima parte biancastre e più larghe alla base che all'apice. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Eritrea, Etiopia. MATERIALE ESAMINATO. Oltre ai due syntypi (vedi sopra), altri 7 esemplari. Eritrea: Asmara, Mai Cuiet, 4.V.1947, 2350 mts., G. de Lotto (3, BMNH); Eritrea, Gura, XII.36-III.37, leg. Nicotra (2, DBAU). Etiopia: Ethiopia, P. Vayssiert (1, MHNP); Ethiopia, Shoa, Lake Langano, V.1989, leg. K. Werner (1, ARCF). 4. Pachytychius aethiopicus n. sp. DIAGNOSI. Un Pachytychius del gruppo aridicola con interstrie elitrali ricoperte da squa- me bianco-grigiastre, poco allungate, disposte in 3-4 serie irregolari, da coricate a subco- ricate, con pronoto fortemente trasverso, con femori posteriori con grosso dente. LOCALITÀ TIPICA. Etiopia, El Dirk. SERIE TIPICA. Holotypus 2: [Etiopia] "El Dirk, 22.5.1939 / Miss. E. Zavattari, Sagan-Omo A.O.l." (MHNP). DESCRIZIONE. 9 (holotypus). Lunghezza mm 4,4. Tegumenti bruno-rossastri ad eccezione di protorace e addome bruno scuri; sul dorso ben visibili fra il rivestimento formato da squame poco allungate (lu/la 2,5-4), subellitti- che, in massima parte biancastre, solo alcune un poco più scure, bruno-grigiastre, sparse sul pronoto e formanti chiazzette scarsamente evidenti sulle elitre, da coricate (in massi- ma parte) a subcoricate, disposte in 3-4 serie molto irregolari per ogni interstria. Strie eli- trali con una serie poco evidente di squame più piccole e strette di quelle delle interstrie. Parte ventrale ricoperta poco fittamente da squame moderatamente allungate (lu/la 4-6). Rostro discretamente più lungo del pronoto (Rlu/Plu 1,20), visto dorsalmente a lati paralleli distintamente divergenti nella parte apicale, visto di lato distintamente e regolar- mente arcuato, debolmente ristretto dall'inserzione delle antenne all'apice. Antenne inseri- te fra terzo medio e terzo apicale; primo segmento del funicolo più robusto e circa 1,5 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto fortemente trasverso (Pla/Plu 1,45), a lati moderatamente arrotondati nel terzo basale, dove è posto il punto più largo, quasi subrettilinei nel terzo medio e ancora distintamente arrotondati nel terzo apicale dove si restringono fortemente, moderatamen- te convesso, con punteggiatura fitta e abbastanza regolare ad eccezione di una stretta stri- scia mediana longitudinale liscia, con intervalli fra i punti stretti e lucidi. Lobi oculari distin- tamente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben visibile. Elitre allungate (Elu/Ela 1,46), a lati subparalleli, con margine anteriore debolmente concavo, con omeri non prominenti in avanti e subarrotondati, lievemente più larghe del pronoto (Ela/Pla 1,12), moderatamente convesse. Interstrie subpiane; strie formate da punti Revisione dei Pachytychius afrotropicali e orientali 143 grossi e profondi, debolmente visibili fra il rivestimento e larghe poco meno delle inter- strie. Femori posteriori con un grosso dente; terzo segmento tarsale bilobato e distintamente più largo del secondo. Spiculum ventrale e spermateca come in P. cognatus (figg. 20 e 25). 3. Non noto. ETIMOLOGIA. Il nome si riferisce alla regione geografica nella quale è stata raccolta la spe- Cie. CONSIDERAZIONI E NOTE COMPARATIVE. Per il robusto dente dei femori posteriori e per le squame dorsali coricate può essere comparato esclusivamente con P. striatellus che, tutta- via, ha rivestimento pit fitto costituito da squame biancastre disposte a strisce sul pronoto e ricoprenti le interstrie elitrali dispari, nonché pronoto meno trasverso. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Specie nota solo della località tipica, situata nell'Etiopia meridionale. MATERIALE ESAMINATO. Nessun altro esemplare oltre all'holotypus (vedi sopra). 5. Pachytychius striatellus n. sp. (figg. 10, 26) DIAGNOSI. Un Pachytychius del gruppo aridicola con interstrie elitrali ricoperte da squa- me che sono di colore bianco su quelle dispari e bruno su quelle pari, poco allungate, dispo- ste in 3-4 serie irregolari, da coricate a subcoricate, con pronoto moderatamente trasverso, con femori posteriori con grosso dente. LOCALITÀ TIPICA. Kenya, Longonot, Nakuru. SERIE TIPICA. Holotypus: 4 " Kenya: Nakuru pr. Longonot, 2000 m, 10.XI.77. Mahnert, Perret" (MHNG). Paratypi: 1 6 e 1 £ con le stesse indicazioni dell'holotypus (MHNG, RCCM); 1 9 [Kenya] "Dr. van Someren, Naivasha, 3.24, V.G.L. van Someren Collection, Brit. Mus. 1959-468" (BMNH). DESCRIZIONE. à (holotypus). Lunghezza mm 2,8. Tegumenti bruno ferruginei, sul dorso quasi completamente nascosti dal fitto rive- stimento formato da squame poco allungate (lu/la 2-4), da ellittiche a sublanceolate, sul pronoto brune e biancastre, queste ultime formanti tre strette strisce, una mediana e due situate verso i lati, sulle elitre grigio-brune e biancastre, le seconde ricoprenti le interstrie dispari così da formare un disegno a strisce, da coricate a subcoricate (queste ultime più scarse e disposte lungo la linea mediana), disposte in 3-4 serie irregolari per ogni interstria. Parte ventrale ricoperta fittamente da squame grigio-brunastre e biancastre, moderatamente allungate (lu/la 3-4), coricate. Rostro un poco più corto del pronoto (Rlu/Plu 0,97), visto dorsalmente a lati paral- leli dalla base all'apice, visto di lato distintamente e regolarmente arcuato, debolmente ristretto dall'inserzione delle antenne all'apice. Antenne inserite fra terzo medio e terzo api- cale; primo segmento del funicolo più robusto e circa 1,3 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,21), a lati molto debolmente arrotonda- ti fin dalla base, con il punto più largo alla metà, debolmente convesso, con punteggiatura 144 CALDARA fitta e abbastanza regolare, con intervalli fra 1 punti stretti e lucidi. Lobi oculari distinta- mente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello di medie dimensioni, ben visibile. Elitre allungate (Elu/Ela 1,45), subrettangolari, con margine anteriore debolmente concavo, con omeri non prominenti in avanti e arrotondati, moderatamente più larghe del pronoto (Ela/Pla 1,24), moderatamente convesse. Interstrie subpiane; strie ben visibili, for- mate da punti profondi e ben visibili, distintamente più strette delle interstrie. Femori posteriori con un distinto dente; terzo segmento tarsale bilobato e distinta- mente più largo del secondo. Edeago come in fig. 10. ? (paratypus). Rostro lievemente più lungo (Rlu/Plu 1,04). Spiculum ventrale come in P. cognatus (fig. 20), spermateca come in fig. 26. VARIABILITÀ. La lunghezza varia da 2,8 a 3,2 mm. Il contrasto fra squame chiare e scure del rivestimento elitrale è più o meno marcato; in due paratipi l'interstria suturale è rico- perta da squame scure. Rlu/Plu 3 0,97-0,99, £ 1,04-1,07; Pla/Plu 1,20-1,24; Elu/Ela 1,43- 1,46; Ela/Pla 1,20-1,24. ETIMOLOGIA. L'aggettivo latino “striatellus” vuole sottolineare la principale caratteristica della specie e cioè il disegno elitrale a strisce. CONSIDERAZIONI E NOTE COMPARATIVE. E' la specie del gruppo con distribuzione più meri- dionale. Dalle altre specie del gruppo considerate in questa sede si distingue facilmente, insieme a P. aethiopicus, per la mancanza di squame sollevate sia sulle elitre che sul pro- noto. Anche da P. aethiopicus, comunque, si può separare senza difficoltà per i caratteri riportati in tabella. Presenta maggiori affinità morfologiche con P. aridicola (Wollaston, 1864), specie largamente diffusa nel Nord Africa (Caldara, 1978, 1984), dal quale differi- sce per la presenza di dente ai femori posteriori ed il rivestimento formato da squame più larghe e più compatte; gli stessi caratteri, oltre alle differenze di rivestimento del pronoto, servono per separare P. striatellus da P. basimaculatus Voss, 1964, la cui diffusione arriva fino al Sudan settentrionale. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Kenya meridionale. MATERIALE ESAMINATO. Solo gli esemplari della serie tipica (vedi sopra). Gruppo di Pachytychius haematocephalus (Gyllenhal) Il gruppo presenta la combinazione dei seguenti caratteri: elitre allungate, da subret- tangolari a debolmente subellittiche, con squame in massima parte moderatamente allun- gate e coricate, solitamente di due colori ben contrastanti fra loro, in ogni caso le più scure ricoprenti gran parte delle prime due interstrie da dove si dipartono a formare fasce e chiaz- ze più o meno numerose e più o meno lunghe; femori posteriori con dente solitamente gros- so; lobo mediano a forma di corto tubo. Al gruppo appartengono una dozzina di specie della Regione Paleartica, cinque spe- cie della regione afrotropicale e le uniche tre specie per ora note della Regione Orientale Revisione dei Pachytychius afrotropicali e orientali 145 (distribuzione in fig. 36). Esso può essere separato dal gruppo di P. aridicola per il diffe- rente tipo di rivestimento elitrale e dal gruppo di P. albosparsus per le elitre allungate (vedi tabella dicotomica). 6. Pachytychius elongatus (Gyllenhal) (figg. 14, 21, 27, 32) Tychius elongatus Gyllenhal, [1835]: 414. Pachytychius elongatus (Gyllenhal), Jekel, 1861: 272. Tournier, 1874: 456. Risbec, 1947: 61. Caldara, 1989: 55. Pachytychius congoanus Hustache, 1923: 20. n. syn. DIAGNOSI. Un Pachytychius del gruppo haematocephalus con pronoto con il punto più largo alla metà da dove si restringe lievemente e gradualmente, in modo subrettilineo, fino in prossimità dell'apice e bruscamente solo all'apice (fig. 32). LOCALITÀ TIPICA. Senegal. SERIE TIPICA. Descritto su esemplari del Senegal, senza più precise indicazioni, dei quali ho esami- nato un d (NHRS) etichettato “Typus / T. elongatus Chevr. / Senegal” (lectotypus qui designato). SINONIMI. Hustache descrive P. congoanus su esemplari raccolti a Fort Lamy (Bas Chari, Chad) senza paragonare la sua nuova specie ad altre del genere. Ho esaminato un synty- pus d di P. congoanus etichettato "Fort Lamy, Congo Français / Pachytychius congoanus m., Hustache det." (MHNP, lectotypus qui designato) perfettamente corrispondente alla descrizione originale. Esso non mostra alcuna differenza, anche nella morfologia degli orga- ni genitali, da P. elongatus. RIDESCRIZIONE. d. Lunghezza mm 5,6. Tegumenti nerastri ad eccezione di rostro, antenne, elitre, tarsi e addome bruno scuri; sul dorso poco visibili fra il fitto rivestimento formato da squame moderatamente allunga- te (lu/la 3-5), subellittiche, coricate, di due colori ben distinti: grigiastre, che sono preva- lenti, e brune. Queste ultime sono più frequenti sul disco del pronoto in sede paramediana, mentre sulle elitre ricoprono una parte prevalente delle prime due interstrie, da dove si dira- mano corte fasce trasversali, e formano alcune piccole chiazze sparse. Parte ventrale rico- perta fittamente da squame bianco-grigiastre e bruno chiare frammiste fra loro, moderata- mente allungate (lu/la 4-6), coricate. Rostro poco più lungo del pronoto (Rlu/Plu 1,03), visto dall'alto a lati paralleli dalla base fino in prossimità dell'apice dove si allargano debolmente, visto di lato distintamen- te e regolarmente arcuato, lievemente ristretto dall'inserzione delle antenne all'apice. Antenne inserite al terzo apicale; primo segmento del funicolo lievemente più grosso e circa 1,5 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,35), a lati moderatamente curvilinei fin dalla base, con il punto più largo alla metà, da dove si restringono lievemente e gradual- mente in modo subrettilineo fino in prossimità dell'apice e bruscamente all'apice (fig. 32), debolmente convesso, con punteggiatura fitta e regolare, con intervalli fra i punti lisci e lucidi. Lobi oculari distintamente pronunciati. Prosterno distintamente incavato nel mezzo. Scutello grande e ben visibile. Elitre allungate (Elu/Ela 1,53), a lati subparalleli nei due terzi anteriori, con margine 146 CALDARA anteriore debolmente concavo, con omeri debolmente prominenti in avanti e arrotondati, lievemente più larghe del pronoto (Ela/Pla 1,11), moderatamente convesse. Interstrie piane; strie poco visibili fra il rivestimento, formate da punti stretti e profondi, distintamente più strette delle interstrie. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Edeago come in fig. 14. 2. Rostro discretamente più lungo (Rlu/Plu 1,18). Spiculum ventrale e spermateca come in 1188.21 27. VARIABILITÀ. Le dimensioni sono comprese fra 5,0 e 6,3 mm (media 5,8 mm, che è anche la misura del lectoytpus). Per quanto riguarda il rivestimento, a volte le squame chiare pre- valgono decisamente anche sulle prime due interstrie elitrali, altre volte le nere formano chiazze molto numerose sulle elitre. Il pronoto ha abbastanza frequentemente il punto di maggior larghezza nel terzo basale. Rlu/Plu & 1,01-1,06, £ 1,13-1,19; Pla/Plu 1,32-1,38; Elu/Ela 1,51-1,59; Ela/Pla 1,09-1,15. CONSIDERAZIONI E NOTE COMPARATIVE. Fra le specie afrotropicali, forma un gruppo morfo- logicamente molto uniforme con P. leucoloma, P. lyali, P. beardae e P. oberprieleri. Da tutte queste, e in particolar modo da P. leucoloma, con il quale sembra convivere in alcu- ne zone (ma verosimilmente su una diversa pianta ospite), differisce solitamente per le maggiori dimensioni (sempre superiori a 5 mm) e per la caratteristica forma del pronoto, poco allargato nella metà apicale e bruscamente ristretto solo all'apice. NOTE BIOLOGICHE. Risbec (1947) descrive il ciclo biologico della specie sulla leguminosa Sesbania aegyptiaca L. L'accoppiamento avviene quando la 9 ha già iniziato da oltre mezz'ora la perforazione del baccello, solitamente in corrispondenza della sua estremità distale. Dopo l'accoppiamento, che dura circa dieci minuti, la 9 riprende il lavoro che la impegna circa per un'altra ora. Una volta completato il foro, essa depone un uovo in pros- simità di un seme, dopo di che ricomincia la stessa operazione a circa 2 cm di ditanza dal buco precedente dirigendosi verso la base del baccello. Dall'uovo, che si presenta ovale, liscio e giallastro ed ha una lunghezza di circa 1 mm, esce una larva di colore avorio che inizia a mangiare il seme più vicino. A maturazione essa misura circa 6-7 mm. Sebbene l'a- dulto sembri preferire nettamente le piante di Sesbania, esso è stato osservato nutrirsi dei baccelli di Indigofera sp. sempre da Risbec, che tuttavia non ha mai trovato uova o larve al loro interno. DISTRIBUZIONE. Africa occidentale, dal Senegal all'Angola. MATERIALE ESAMINATO. Oltre agli esemplari delle serie tipiche (vedi sopra), altri 33 esem- plari. Senegal: Senegal, coll. Jekel (1, MSNM); Senegal, ex Desbrochers (1, MSNM); Sénégal, Coll. Roelofs (1, ISNB); Sénégal, Coll. Dejean (4, ISNB); Sénégal, Coll. Castelnau (1, ISNB); Senegal, Bambey, 30.X.1940, J. Risbec, on millet (2, BMNH); idem eccetto 1946 (2, BMNH); idem eccetto 15.1.1945, on Sesbania aegyptiaca (7, BMNH); Bambey, Coll. Jekel (2, MSNM); Senegal, Sébikotane, 20/1/1946, H. Durand (4, MHNP). Nigeria: Kano, N. Nigeria, Nov. 1951, W. E. S. Merrett (1, BMNH). Gabon: Gabon, coll. Jekel (1, MSNM). Angola: Angola, Luanda, 20.v111.1949, G. R. Gradwell & D. Snow (1, BMNH); Angola, 76.28 (3, BMNH). Senza indicazioni (2, BMNH). Revisione dei Pachytychius afrotropicali e orientali 147 n; 05 ; Figg. 7:42 Edeago di: 7 — Pachytychius cognatus n. sp.; 8 — P. setosus Marshall; 9 DO. erythreensis Hustache; 10 — P. striatellus n. sp.; 11 — P. albosparsus (Fähraeus); 12 — P. tuberculatus n. sp.; 13 — P. viciae Marshall. Scala = mm 0,50. 148 CALDARA 7. Pachytychius leucoloma Jekel (figg. 3, 15, 31) Pachytychius leucoloma Jekel, 1861: 272. DIAGNOSI. Un Pachytychius del gruppo haematocephalus con pronoto moderatamente tra- sverso, a lati moderatamente arrotondati fin dalla base, con il punto più largo nel terzo ante- riore da dove si restringe nettamente fino all'apice (fig. 31), moderatamente convesso. LOCALITÀ TIPICA. Senegal. SERIE TIPICA. Specie descritta sinteticamente da Jekel su esemplari del Senegal (coll. Dejean), dei quali ho esaminato una © (ISNB) etichettata "Sénégal / Coll. Dejean, Coll. Roelofs / Tychius leuco- loma m., L. in Senegal, D. Damosin / Pachytychius leucoloma Jekel / Type" (lectotypus qui desi- gnato). RIDESCRIZIONE. 6. Lunghezza mm 4,2. Tegumenti nerastri ad eccezione di rostro, antenne, elitre, tarsi e addome bruno scuri; sul dorso poco visibili fra il fitto rivestimento formato da squame poco allungate (lu/la 2- 4), da subellittiche a subovali, coricate, di due colori ben contrastanti: biancastre e bruno scure, queste ultime ricoprono interamente il disco del pronoto e quasi completamente le prime e le ultime due interstrie delle elitre dove formano anche alcune fasce trasverse, che vanno dalla sutura alla quarta interstria, e alcune chiazzette sparse. Parte ventrale ricoper- ta fittamente da squame bianco-grigiastre e bruno chiare frammiste fra loro, moderatamente allungate (lu/la 4-6), coricate. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,12), visto dall'alto un poco allargato nella parte basale e in quella apicale dall'inserzione delle antenne all'apice, visto di lato distintamente e regolarmente arcuato, lievemente ristretto dall'inserzione delle anten- ne all'apice. Antenne inserite fra terzo medio e terzo apicale; primo segmento del funico- lo antennale lievemente più grosso e quasi della stessa lunghezza del secondo. Fronte un poco più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,36), a lati moderatamente arrotondati fin dalla base, con il punto più largo nel terzo anteriore da dove si restringe nettamente fino all'apice (fig. 31), moderatamente convesso, con punteggiatura piccola e regolare che rispar- mia una stretta striscia mediana longitudinale, con intervalli fra i punti lisci e lucidi. Lobi oculari distintamente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,57), a lati paralleli nei due terzi anteriori, con margine ante- riore moderatamente concavo, con omeri debolmente prominenti in avanti, un poco più lar- ghe del pronoto (Ela/Pla 1,14), moderatamente convesse. Interstrie piane; strie poco visi- bili fra il rivestimento, formate da punti stretti e profondi, distintamente più strette delle interstrie. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Fdeago come in fig. 15. 2. Rostro un poco più lungo (Rlu/Plu 1,17). Spiculum ventrale e spermateca come in P. elongatus (figg. 21 e 27). VARIABILITÀ. Le dimensioni variano da 3,5 a 6,0 mm (il lectotypus misura mm 4,1), seb- bene gli esemplari sopra 1 5 mm siano poco frequenti. Per quanto riguarda il rivestimento, Revisione dei Pachytychius afrotropicali e orientali 149 le fasce e le chiazze elitrali possono essere più o meno numerose; a volte una fascia fra terzo medio e terzo posteriore delle elitre congiunge le squame scure delle prime due inter- strie a quelle delle ultime. Il pronoto può avere lati solo debolmente curvilinei e più distin- tamente divergenti dalla base al terzo apicale. Rlu/Plu & 1,07-1,13, 2 1,15-1,20; Pla/Plu 1,32-1,45; Elu/Ela 1,54-1,59; Ela/Pla 1,10-1,16. CONSIDERAZIONI E NOTE COMPARATIVE. In tutte le collezioni da me esaminate, P. leucoloma è sempre stato confuso con P. elongatus, insieme al quale sembra trovarsi in alcune loca- lità del Senegal, probabilmente su una diversa leguminosa, sebbene non possa escludersi, in base ai dati disponibili (esemplari raccolti da Risbec), che le due specie vivano entram- be sulla stessa pianta. A parte le differenze nella struttura del lobo mediano (in P. leucolo- ma l'endofallo è sprovvisto di scleriti subapicali), P. leucoloma è solitamente più piccolo (solo tre esemplari di quelli da me esaminati rientrano nell’ambito di lunghezza di P. elon- gatus) e soprattutto ha pronoto più quadrato esssendo il punto più largo situato nella metà anteriore: inoltre sul disco del pronoto le squame chiare sono molto scarse, mentre in P. elongatus solitamente queste risultano numerose. NOTE BIOLOGICHE. Come risulta dai cartellini degli esemplari raccolti da Risbec, la pianta ospite potrebbe essere Vigna sp.(Fabaceae); tuttavia, sempre dalle raccolte di Risbec e visto che le specie di Pachytychius sono di rado strettamente monofaghe, non può escludersi che la specie possa svolgere il suo ciclo biologico anche su Sesbania aegyptiaca, come P. elon- gatus (vedi note biologiche di questa specie), oppure nutrirsi dei tessuti di questa pianta solamente allo stato adulto. DISTRIBUZIONE. Senegal, Nigeria, Alto Volta, Chad. MATERIALE ESAMINATO. Oltre al lectotypus (vedi sopra), altri 19 esemplari. Senegal: Senegal, Bambey, 15.1.1945, J. Risbec, on Vigna (9, BMNH); idem, on Sesbania aegyptiaca (2, BMNH). Nigeria: Exped. Mus. G. Frey, Nigeria-Kamerun, Bechyne 1955-56 / Kano, 2.X.55 (1, BMNH); N. Nigeria, Badeggi, Sep. 1910, J. W. Scott-Macfie, 1911-417 (1, BMNH); Bantschi to Lokoja, N. Nigeria, L. M. Bucknill, 1908-229 (1, BMNH). Alto Volta: Africa, Upper Volta, Pabrè near Quagadougou, R. P. Fernandez (4, CWOB). Chad: Coll. Mus. Congo, Tchad: iles du lac, XI/XII.1957, P. Renaud (1, MACT). 8. Pachytychius lyali n. sp. (16, 34) DIAGNOSI. Un Pachytychius del gruppo haematocephalus con interstrie elitrali, soprattut- to nel terzo apicale, con una serie mediana irregolare di squame lievemente sollevate dai tegumenti, visibili guardando le elitre di profilo; con pronoto fortemente trasverso, a lati fortemente arrotondati fin dalla base, con il punto più largo alla metà (fig. 34), distinta- mente convesso. LOCALITÀ TIPICA. Zambia, Mwengwa. SERIE TIPICA. Holotypus d: [Zambia] "N. W. Rhodesia: Mwengwa 27°40'E. 13°S, 3.IV.1914, H. C. Dollman / H. C. Dollman Coll. 1919-79" (BMNH) (manca la zampa anteriore sinistra); | paratypus ® con le stesse indicazioni dell'holotypus eccetto "4.IV.1914" (RCCM). DESCRIZIONE. 6 (holotypus). Lunghezza mm 4,4. Tegumenti nerastri ad eccezione di rostro, antenne e zampe bruno scuri; sul dorso 150 CALDARA scarsamente visibili fra il fitto rivestimento formato da squame poco allungate (lu/la 3-4), subellittiche, di due colori ben contrastanti: biancastre e bruno-nerastre. Queste ultime sono più abbondanti sul disco del pronoto e sulla prima interstria elitrale e formano, sempre sulle elitre, numerose strette fasce trasverse, che vanno dalla sutura verso 1 lati, e alcune picco- le chiazze sparse. Le squame sono tutte coricate ad eccezione di alcune lievemente solle- vate sulle interstrie elitrali, soprattutto nel terzo apicale; queste ultime sono ben visibili guardando le elitre di profilo e formano una serie mediana irregolare su ogni interstria. Parte ventrale ricoperta fittamente da squame bianco-grigiastre e bruno chiare frammiste fra loro, discretamente allungate (lu/la 5-8), coricate. Rostro lungo circa quanto il pronoto (Rlu/Plu 0,99), visto dall'alto a lati debolmente convergenti dalla base all'apice, visto di lato moderatamente e regolarmente arcuato, lie- vemente ristretto dalla base all'apice. Antenne inserite al terzo apicale; primo segmento del funicolo più grosso e circa 1,7 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto fortemente trasverso (Pla/Plu 1,46), a lati fortemente arrotondati fin dalla base, con il punto più largo alla metà (fig. 34), distintamente convesso; con punteggiatura fitta, abbastanza regolare, con intervalli fra 1 punti stretti e lucidi. Lobi oculari moderata- mente pronunciati. Prosterno con margine anteriore fortemente incavato. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,44), a lati subparalleli nei due terzi anteriori, con margine anteriore moderatamente concavo, con omeri debolmente prominenti in avanti, un poco più larghe del pronoto (Ela/Pla 1,14), moderatamente convesse. Interstrie piane; strie scar- samente visibili fra il rivestimento, formate da punti piccoli e profondi, distintamente più strette delle interstrie. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Edeago come in fig. 16. 2 (paratypus). Come il 4 ad eccezione di rostro discretamente più lungo (Rlu/Plu 1,10). Spiculum ventrale e spermateca come in P. elongatus (figg. 21 e 27). VARIABILITA. Il paratypus ha dimensioni maggiori rispetto all’ holotypus (mm 4,9). ETIMOLOGIA. La specie è dedicata a Chris Lyal in segno di ringraziamento per l'indispen- sabile aiuto fornitomi durante le mie visite al BMNH. CONSIDERAZIONI E NOTE COMPARATIVE. Specie strettamente affine soprattutto a P. ober- prieleri e P. beardae. Oltre che per la forma del lobo mediano, differisce da entrambi per il pronoto più trasverso e solo poco più largo delle elitre, più convesso sul disco e a lati più arrotondati, e per la serie di squame lievemente sollevate sulle interstrie elitrali. Da P. bear- dae differisce, oltre che per questi particolari, anche per 1 tegumenti in gran parte nerastri e per le squame del dorso di colori molto più contrastanti fra loro (biancastre e bruno-nera- stre invece di grigio-biancastre e bruno chiare). NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Zambia occidentale. MATERIALE ESAMINATO. Solo gli esemplari della serie tipica (vedi sopra). Revisione dei Pachytychius afrotropicali e orientali Foul 9. Pachytychius beardae n. sp. (figg. 18, 33) DiaGNOSI. Un Pachytychius del gruppo haematocephalus con disco del pronoto e prime due interstrie elitrali ricoperti da squame circa dello stesso colore delle altre; con tegumenti completamente bruno-rossastri; con pronoto distintamente trasverso, gradualmente ristret- to nel terzo apicale (fig. 33), debolmente convesso. LOCALITA TIPICA. Zambia, Namwala. SERIE TIPICA. Holotypus d : [Zambia] "N. W. Rhodesia: Namwala. 12.V1.1914, H. C. Dollman. / Dry grass-roots nr. R. Kaful / H. C. Dollman Coll. 1919-79" (BMNH); 3 paratypi con le stesse indica- zioni dell'holotypus (2 9 2 BMNH, 1 4 RCCM). DESCRIZIONE. d (holotypus). Lunghezza mm 4,3. Tegumenti completamente bruno-rossastri; sul dorso un poco visibili fra il rivesti- mento formato da squame abbastanza fitte, poco allungate (lu/la 3-5), subellittiche, cori- cate, di due colori poco contrastanti fra loro: grigio-biancastre e bruno chiare; queste ulti- me sono più frequenti sul disco del pronoto, mentre sulle elitre formano piccole chiazze sparse. Parte ventrale ricoperta abbastanza fittamente da squame biancastre, moderatamente allungate (lu/la 4-6), coricate. Rostro lievemente più lungo del pronoto (Rlu/Plu 1,04), visto dall'alto a lati paralle- li, visto di lato moderatamente e regolarmente arcuato, lievemente allargato ventralmente alla metà (margine ventrale lievemente sinuoso), un poco ristretto dall'inserzione delle antenne all'apice. Antenne inserite nel terzo apicale; primo segmento del funicolo più gros- so e circa 1,8 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. Pronoto distintamente trasverso (Pla/Plu 1,40), a lati moderatamente curvilinei fin dalla base, con il punto più largo alla metà (fig. 33), debolmente convesso, con punteggia- tura fitta, abbastanza regolare, con intervalli fra i punti stretti e lucidi. Lobi oculari mode- ratamente pronunciati. Prosterno con margine anteriore moderatamente incavato. Scutello grande, ben visibile. Elitre allungate (Elu/Ela 1,45), a lati subrettilinei nei due terzi anteriori, con margi- ne anteriore debolmente concavo, con omeri debolmente prominenti in avanti, discreta- mente più larghe del pronoto (Ela/Pla 1,20), moderatamente convesse. Interstrie piane; strie scarsamente evidenti fra il rivestimento, formate da punti piccoli e profondi, distintamen- te più strette delle interstrie. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Edeago come in fig. 18. ? (paratypus). Rostro un poco più lungo (Rlu/Plu 1,08). Spiculum ventrale e sper- mateca come in P. elongatus (figg. 21 e 27). VARIABILITA. Le dimensioni variano da 4,2 a 4,5 mm. Per il resto non si evidenziano varia- zioni degne di nota fra gli esemplari della serie tipica. Rlu/Plu 4 1,02-1,04, 2 1,07-1,08; Pla/Plu 1,39-1,44; Elu/Ela 1,43-1,46; Ela/Pla 1,16-1,21. ETIMOLOGIA. Dedico con piacere la specie a Jane Beard in segno di ringraziamento per l'aiu- to fornitomi nell'esame delle collezioni entomologiche del BMNH. CONSIDERAZIONI E NOTE COMPARATIVE. Si distingue dalle specie vicine, in primo luogo P. 152 CALDARA oberprieleri e P. lyali, per i tegumenti completamente rossastri sia sul dorso che ventral- mente. Oltre che per la forma del lobo mediano, differisce da entrambe queste specie per le squame dorsali di due colori scarsamente contrastanti fra loro, e da P. lyali anche per il pronoto meno largo rispetto alle elitre, meno convesso sul disco e a lati meno arrotondati. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Zambia occidentale. MATERIALE ESAMINATO. Solo gli esemplari della serie tipica (vedi sopra). 10. Pachytychius oberprieleri n. sp. (fig. 17) DIAGNOSI. Un Pachytychius del gruppo haematocephalus con pronoto distintamente tra- sverso, a lati moderatamente curvilinei fin dalla base, con il punto più largo alla metà, mode- ratamente convesso. LOCALITÀ TIPICA. Sud Africa, Natal, Zululand, Empangeni. SERIE TIPICA. Holotypus à : [Sud Africa] "Empangeni, Zululand, 7.3.67, E. Borman" (SANC). 3 paratypi 9 2: 1 "South Africa, Natal, Empangeni, 28.44S 31.54E, 17.11.1984, P. Reavell" (RCMC); 1 [Sud Africa] "Zululand, Lr. Umhlatuzi R., 6.vi.1926 / S. Africa, R. E. Turner, Brit. Mus. 1926-277" (BMNH); 1 [Sud Africa, Transvaal] "Plat River, 6-18.4.05, Waterberg Dis., C. Swierstra" (SANC). DESCRIZIONE. 3 (holotypus). Lunghezza mm 4,3. Tegumenti bruno-rossastri ad eccezione della parte laterale del disco del pronoto, delle prime due e delle ultime due interstrie elitrali e dei lati della parte inferiore nerastri; sul dorso un poco visibili fra il rivestimento abbastanza fitto formato da squame poco allun- gate (lu/la 3-5), subellittiche, coricate, in massima parte bruno-grigiastre, e bruno più scure ricoprenti sulle elitre parte delle prime due interstrie e formanti piccole chiazze sparse sul disco del pronoto. Parte ventrale ricoperta abbastanza fittamente da squame bianco-gri- giastre e bruno chiare, discretamente allungate (lu/la 4-7), coricate. Rostro lievemente più lungo del pronoto (Rlu/Plu 1,07), visto dall'alto a lati subpa- ralleli, visto di lato non ristretto e moderatamente e regolarmente arcuato dalla base all'a- pice. Antenne inserite fra terzo medio e terzo apicale; primo segmento del funicolo un poco più robusto e circa 1,5 volte più lungo del secondo. Fronte un poco più stretta del rostro alla base. | Pronoto distintamente trasverso (Pla/Plu 1,41), a lati moderatamente curvilinei fin dalla base, con il punto più largo alla metà, moderatamente convesso, con punteggiatura fitta, abbastanza regolare, con intervalli fra i punti stretti e lucidi. Lobi oculari distinta- mente pronunciati. Prosterno con margine anteriore distintamente incavato. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,47), a lati subparalleli nei tre quarti basali, con margine anteriore debolmente concavo, con omeri debolmente prominenti in avanti, discretamente più larghe del pronoto (Ela/Pla 1,25), moderatamente convesse. Interstrie debolmente con- vesse; strie poco visibili, con punti poco profondi, distintamente più strette delle interstrie. Femori posteriori con forte dente aguzzo; terzo segmento tarsale bilobato e distinta- mente più largo del secondo. Edeago come in fig. 17. Revisione dei Pachytychius afrotropicali e orientali 153 a 19 | 18 Figg. 14-19. Edeago di: 14 — Pachytychius elongatus (Gyllenhal); 15 — P. leucoloma Jekel; 16 - P. lyali n. sp.; 17 - P. oberprieleri n. sp.; 18 — P. beardae n. sp.; 19 — P. indicus Tournier. Scala = mm 0,50. 154 CALDARA ? (paratypus). Rostro un poco più lungo (Rlu/Plu 1,11). ER ventrale e sper- mateca come in P. elongatus (figg. 21 e 27). VARIABILITÀ. La lunghezza è compresa fra 4,0 e 4,8 mm. Non si rilevano altre differenze degne di nota fra gli esemplari della serie tipica. Rlu/Plu £ 1,10-1,15; Pla/Plu 1,39-1,42; Elu/Ela 1,45-1,50; Ela/Pla 1,22-1,25. ETIMOLOGIA. Dedico con piacere la specie a Rolf Oberprieler che mi ha inviato in studio numerosi e interessanti esemplari indispensabili per questa mia revisione. CONSIDERAZIONI E NOTE COMPARATIVE. Specie strettamente affine a P. beardae, dal quale differisce solamente per i tegumenti di protorace e addome bruno nerastri e non rossastri, per le squame scure del rivestimento dorsale più marcatamente contrastate dalle chiare e per la forma del lobo mediano che appare più corto e allargato che in tutte le altre specie ad esso correlate (P. elongatus, P. lyali e P. leucoloma). NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Sud Africa (Natal, Transvaal). MATERIALE ESAMINATO. Solo gli esemplari della serie tipica (vedi sopra). 11. Pachytychius indicus Tournier (fig. 19) Pachytychius indicus Tournier, 1874: 454, nota. DrAGNOSI. Un Pachytychius del gruppo haematocephalus con pronoto distintamente tra- sverso, a lati debolmente arrotondati nella metà basale, più marcatamente nella metà api- cale, restringendosi bruscamente in prossimità dell'apice, con il punto più largo un poco oltre la metà, subpiano. LOCALITÀ TIPICA. India, Bombay. SERIE TIPICA. Specie descritta su esemplari di Bombay già in precedenza classificati da Jekel con tale nome rimasto però in litteris. Mentre nella collezione Tournier non sono riuscito a trovare esempla- ri di P. indicus, fra il materiale della collezione Bowring (BMNH) ho reperito un d (mancante di zampa media e posteriore sinistra e con 1 due terzi apicali dell’elitra destra staccata dalla parte restan- te del corpo) etichettato "Bombay / Boly / Indicus (Jekel) Bombay / Bowring 63-47" ben corrispon- dente alla seppur breve e generica descrizione di Tournier (lectotypus qui designato). E' tuttavia da notare che in collezione Solari (MSNM) ho trovato due esemplari provenienti dalla collezione Jekel ed etichettati "Bombay, coll. Jekel (scritto da Solari) / nov. sp. indicus Jekel / Pachytychius indicus (Jek.) Bombay / indicus Tourn. typus! ex coll. Jekel (ancora scritto da Solari)". In realtà questi due esemplari non risultano differenti dai comuni P. elongatus del Senegal e non li considero pertanto appartenenti alla serie tipica di P. indicus. E’ probabile che un errore di cartelli- natura abbia tratto in inganno Solari. E' interessante, infatti, sottolineare la curiosa coincidenza che P. elongatus è stato trovato da più raccoglitori a Bambay, una località del Senegal quasi omonima della città indiana Bombay. RIDESCRIZIONE. é. Lunghezza mm 4,7. Tegumenti bruno-nerastri ad eccezione di rostro, antenne, tibie e tarsi di colore bruno ferrugineo scuro, sul dorso poco visibili fra il fitto rivestimento formato da squame poco allungate (lu/la 3-5), da subellittiche a sublanceolate, coricate, brune e bianco-giallastre, formanti un disegno marmorizzato sia sul pronoto che sulle elitre dove le scure ricoprono Revisione dei Pachytychius afrotropicali e orientali 155 in massima parte le prime due interstrie. Parte ventrale ricoperta fittamente da squame bian- co-grigiastre e bruno chiare, discretamente allungate (lu/la 4-8), coricate. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,12), visto dall'alto a lati sub- paralleli, visto di lato fortemente e regolarmente arcuato, lievemente ristretto dall'inser- zione delle antenne all'apice. Antenne inserite nel terzo apicale; primo segmento del funi- colo un poco più grosso e circa 1,5 volte più lungo del secondo. Fronte distintamente più stretta del rostro alla base. Pronoto distintamente trasverso (Pla/Plu 1,42), a lati debolmente arrotondati nella metà basale, più marcatamente nella metà apicale, restringendosi bruscamente in prossi- mità dell'apice, con il punto più largo un poco oltre la metà, debolmente convesso, con pun- teggiatura piccola e regolare, con intervalli fra 1 punti stretti e lucidi. Lobi oculari distinta- mente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,54), a lati molto debolmente curvilinei nei due terzi ante- riori, con margine anteriore moderatamente concavo, con omeri debolmente prominenti anteriormente, moderatamente più larghe del pronoto (Ela/Pla 1,14), moderatamente con- vesse. Interstrie piane; strie scarsamente visibili fra il rivestimento, formate da punti allun- gati e profondi, larghe come la meta della larghezza di un'interstria. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Edeago come in fig. 19. 2. Rostro moderatamente più lungo (Rlu/ Plu 1,23). Spiculum ventrale e spermate- ca come in P. elongatus (figg. 21 e 27). VARIABILITÀ. La lunghezza varia da 4,4 (misura anche del lectotypus) a 5,0 mm. Un d da me esaminato presenta una evidente malformazione del rostro, che è abnormemente corto e fortemente ricurvo. Rlu/Plu & 1,08-1,13, 9 1,18-1,24; Pla/Plu 1,40-1,45; Elu/Ela 1,52- 1,55; Ela/Pla 1,10-1,15. CONSIDERAZIONI E NOTE COMPARATIVE. Specie estremamente simile a P mungonis, dal quale sembra differire solo per il pronoto più trasverso e rivestito da squame prevalentemente chiare. Queste esili differenze, insieme alla diversa pianta ospite, mi hanno convinto a tene- re separate per il momento le due specie. Pachytychius indicus risulta decisamente simile anche a P. elongatus, dal quale differisce essenzialmente per il pronoto un poco più tra- sverso e arrotondato. NOTE BIOLOGICHE. La pianta ospite è verosimilmente Sesbania aculeata sulla quale è stato raccolto un adulto, mentre dai suoi semi è uscito un altro esemplare da me esaminato ("dain- cha" è infatti il nome indiano dato a S. aculeata, Lyal com. pers.). E' da sottolineare che sullo stesso genere di leguminose vive in Africa la specie affine P. elongatus. DISTRIBUZIONE. India centro-meridionale. MATERIALE ESAMINATO. Oltre al lectotypus, altri 6 esemplari. India: Bombay (2, BMNH); Bombay, E. M. Janson (1, BMNH); India, Bombay, Karjat, 29.X.1955, S. K. Dorge, on Sesbania aculeata" (1, BMNH); South India, Coimbatore, 28.11.1916 (2, BMNH). 156 CALDARA 12. Pachytychius mungonis Marshall Pachytychius mungonis Marshall, 1915: 379. DIAGNOSI. Un Pachytychius del gruppo haematocephalus con pronoto moderatamente tra- sverso, a lati moderatamente arrotondati, con il punto più largo alla metà, subpiano. LOCALITÀ TIPICA. India, Madras. SERIE TIPICA. Specie descritta su esemplari raccolti nell'antico stato coloniale di Madras nell'India meridionale (distretti di Bellary e di Kurnul) senza stabilire l'holotypus. AI BMNH ho esaminato un 6 etichettato " Type / Madras 1915-116 / XI.11, on green-gram pods, Hadagalli Madras YR / Pachytychius mungonis Mshl. Type" (lectotypus qui designato) e due £ 2 "Cotype / Bellary Dt., Hadagalli, 5-9 Dec. 12 / G. A. K. Marshall Coll., B. M. 1950-255". Tutti i tre esemplari hanno perso parte delle squame del rivestimento, mentre al lectotypus manca la zampa posteriore sinistra. RIDESCRIZIONE. 6. Lunghezza mm 4,2. Tegumenti bruno scuri ad eccezione di antenne e terzo segmento tarsale più chiari e protorace ed interstria suturale elitrale bruno-nerastri, sul dorso poco visibili fra il fitto rive- stimento formato da squame poco allungate (lu/la 2-4), da subellittiche a sublanceolate, coricate, bruno scure e bianco-grigiastre. Sul pronoto le squame bruno scure sono decisa- mente prevalenti, mentre le biancastre formano una stretta striscia longitudinale mediana e chiazzette confuse ai lati e fra lati e linea mediana. Sulle elitre le più scure ricoprono le prime due interstrie e formano numerose chiazzette sparse sulla restante porzione. Parte ventrale ricoperta fittamente da squame biancastre e grigiastre, poco allungate (Elu/Ela 1,5- 3), e da alcune squame lunghe setoliformi, biancastre. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,12), visto dall'alto a lati sub- paralleli, visto di lato fortemente e regolarmente arcuato, lievemente ristretto dall'inser- zione delle antenne all'apice. Antenne inserite nel terzo apicale; primo segmento del funi- colo un poco più grosso e circa 1,5 volte più lungo del secondo. Fronte distintamente più stretta del rostro alla base. Pronoto moderatamente trasverso (Pla/Plu 1,33), a lati moderatamente arrotondati, con il punto più largo alla metà, debolmente convesso, con punteggiatura piccola e rego- lare, con intervalli fra 1 punti stretti e lucidi. Lobi oculari distintamente pronunciati. Prosterno moderatamente incavato nel mezzo. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,49), a lati molto debolmente curvilinei nei due terzi ante- riori, con margine anteriore moderatamente concavo, con omeri debolmente prominenti anteriormente, moderatamente più larghe del pronoto (Ela/Pla 1,17), moderatamente con- vesse. Interstrie piane; strie scarsamente visibili fra il rivestimento, formate da punti allun- gati e profondi, larghe come la metà della larghezza dell'interstria. Femori posteriori con grosso e aguzzo dente; terzo segmento tarsale bilobato e distin- tamente più largo del secondo. Edeago come in P. indicus (fig. 19). 2. Rostro un poco più lungo (Rlu/Plu 1,18). Spiculum ventrale e spermateca come in P. elongatus (figg. 21 e 27). VARIABILITÀ. La lunghezza varia da 3,9 a 4,8 mm (il lectotypus misura mm 4,3). Sul pro- noto le chiazzette di squame chiare, sempre poco numerose, risultano più o meno contra- Revisione dei Pachytychius afrotropicali e orientali ha 31 32 25 29 SO 33 34 Figg. 20-34. Spiculum ventrale (20-24), spermateca (25-30) e-pronoto (31-34) di: 20 — Pachytychius cognatus n. sp.; 21 — P. elongatus (Gyllenhal); 22 — P. viciae Marshall; 23 — P. tuberculatus n. sp.; 24 — P. albosparsus (Fahraeus); 25 — P. cognatus n. sp.; 26 — P. striatellus n. sp.; 27 — P. elongatus (Gyllenhal); 28 — P. viciae Marshall; 29 — P. tuberculatus n. sp.; 30 - P. albosparsus (Fahraeus); 31 — P. leucoloma Jekel; 32 — P. elongatus (Gyllenhal); 33 — P. beardae n. sp.; 34 — P. lyali n. sp. Scala = mm 0,25 (figg. 20-30); mm 0,50 (figg. 31-34). 158 CALDARA state rispetto alle squame scure. Rlu/Plu d 1,09-1,13, 2 1,17-1,20; Pla/Plu 1,30-1,35; Elu/Ela 1,46-1,50; Ela/Pla 1,16-1,20. CONSIDERAZIONI E NOTE COMPARATIVE. Specie strettamente imparentata con P. indicus. Per le differenze vedi note comparative di quest'ultima specie e tabella dicotomica. NOTE BIOLOGICHE. Secondo le osservazioni effettuate da Bainbrigge Fletcher (Marshall, 1915), la specie parassita Phaseolus mungo L., spesso causando notevoli danni alle coltu- re, Nei semi di questa leguminosa la £ depone le uova dopo aver prodotto un foro nel bac- cello. Solitamente si trovano tre uova per ogni seme, due ai lati e una nel mezzo. La larva si nutre di 3-4 semi prima di arrivare a maturazione: a questo punto emerge dal baccello e si lascia cadere nel terreno dove avviene la trasformazione a pupa. DISTRIBUZIONE. India centro-meridionale. MATERIALE ESAMINATO. Oltre agli esemplari della serie tipica (vedi sopra), altri 4 esemplari. India: Deccan, Ahmadnagar, 1932, J. E. M. Boyd, B. M. 1934-37 (1, BMNH); Godavari Dt., Thanelanka, 22-24 Dec. 12, T.V.R. coll., on Red gram (1, BMNH); S. India, Campbell (2, BMNH). 13. Pachytychius viciae Marshall (figg. 4, 13, 22, 28) Pachytychius viciae Marshall, 1938: 160. DIAGNOSI. Un Pachytychius del gruppo haematocephalus con le squame più scure rico- prenti sulle elitre tutt'al più le prime due interstrie e formanti solo qualche confusa chiaz- za sulle altre interstrie, con pronoto debolmente trasverso, con femori posteriori con pic- colo dente. LOCALITÀ TIPICA. India, Uttar Pradesh, Dehra Dun. SERIE TIPICA. Specie descritta su esemplari raccolti a Dehra Dun nell'Uttar Pradesh (India settentrio- nale), dei quali ne ho esaminati nove (BMNH), uno dei quali etichettato "Dehra Dun, U.P., J.C.M. Gardner, 2.V.1936 / on Vicia sativa pods / Pres. by Imp. Inst. Ent. B.M. 1936-714 / Pachytychius viciae Mshl., Type male / Type" (lectotypus qui designato), mentre gli altri otto (4 88 e 4 £ 2) por- tano identiche indicazioni ad eccezione di un cartellino con scritto "Cotype" (2) o "Paratype" (6). RIDESCRIZIONE. d (lectotypus). Lunghezza mm 3,0. Tegumenti bruno-rossastri; sul dorso pressoché completamente nascosti dal fitto rive- stimento formato da squame in parte embricate, di forma lanceolata, coricate, biancastre e bruno chiare con evidenti riflessi sericei. Sul pronoto le squame sono larghe (lu/la 1,5-2), mentre sulle interstrie elitrali sono un poco più lunghe (lu/la 2-3) soprattutto alcune che tendono a formare una serie mediana e che sono debolmente sollevate dai tegumenti. Le squame brune ricoprono prevalentemente il disco del pronoto e le prime due interstrie eli- trali. Strie elitrali scarsamente visibili, con una serie di strette e lunghe squame biancastre. Anche la parte ventrale ricoperta fittamente da squame biancastre, di forma subellittica o subovale, larghe (lu/la 2-4), in mezzo alle quali sono frammiste sugli ultimi sterniti alcu- ne lunghe squame setoliformi un poco sollevate (lu/la 5-6). Rostro poco più lungo del pronoto (Rlu/Plu 1,04), visto dall'alto a lati subparalleli, visto di lato moderatamente arcuato, lievemente ristretto dall'inserzione delle antenne all'a- pice. Antenne inserite fra terzo medio e terzo apicale; primo segmento del funicolo un poco Revisione dei Pachytychius afrotropicali e orientali cognatus setosus erythreensis aethiopicus striatellus albosparsus tuberculatus 159 Fig. 35. Areale di diffusione dei gruppi di P. aridicola (/////) e di P. albosparsus (\\\\\) e località det- tagliate di raccolta delle specie afrotropicali. 160 CALDARA più grosso e circa 1,5 volte più lungo del secondo. Fronte debolmente più stretta del rostro alla base. Pronoto debolmente trasverso (Pla/Plu 1,19), a lati un poco curvilinei e divergenti dalla base al terzo apicale dove è situato ıl punto più largo, debolmente convesso, con pun- teggiatura piccola e regolare, con intervalli fra 1 punti stretti e lucidi (visibili solamente negli esemplari mal conservati). Lobi oculari moderatamente pronunciati. Prosterno mode- ratamente incavato nel mezzo. Scutello grande, ben evidente. Elitre allungate (Elu/Ela 1,60), con margine anteriore moderatamente concavo, con omeri non prominenti in avanti e arrotondati, discretamente più larghe del pronoto (Ela/Pla 1,27), moderatamente convesse. Interstrie piane; strie scarsamente visibili perché nascoste dal rivestimento, formate da punti allungati e profondi, apparentemente molto più strette dell’interstria. Femori posteriori con un piccolo ma distinto dente; terzo segmento tarsale bilobato e moderatamente ma distintamente più largo del secondo. Edeago come in fig. 13. ? (paralectotypus). Rostro moderatamente più lungo (Rlu/Plu 1,10). Spiculum ven- trale e spermateca come in figg. 22 e 28. VARIABILITÀ. La lunghezza varia da 2,6 a 3,2 mm. Per quanto riguarda il rivestimento, sul pronoto le squame più chiare possono essere numerose anche sul disco e non solo ai lati; viceversa, in un esemplare le squame scure prevalgono sulle chiare che formano una stret- ta stria centrale e due strie per ogni lato. Sulle elitre le squame brune, oltre alle prime due interstrie, possono ricoprire in minima parte anche le altre interstrie; in alcuni esemplari sulle interstrie non è evidente la serie centrale di rade squame un poco sollevate. Infine, il pronoto può presentare lati rettilinei nei due terzi basali. Rlu/Plu & 1,02-1,05, 9 1,08-1,13; Pla/Plu 1,15-1,20; Elu/Ela 1,56-1,63; Ela/Pla 1,26-1,32. CONSIDERAZIONI E NOTE COMPARATIVE. Risulta facilmente distinguibile dalle altre due spe- cie per il momento note del subcontinente indiano soprattutto per le minori dimensioni del corpo e del dente dei femori posteriori. NOTE BIOLOGICHE. Come riportato sul cartellino degli esemplari della serie tipica, la spe- cie risulta raccolta su legumi di Vicia sativa L. che, verosimilmente, costituisce la pianta o una delle piante ospiti. Per quanto riguarda l’esemplare raccolto in Pakistan su melograno (Punica granata L., Punicaceae), è probabile che tale pianta sia solo una pianta di rifugio. DISTRIBUZIONE. India settentrionale, Pakistan orientale. MATERIALE ESAMINATO. Oltre ai 9 esemplari della serie tipica solo altri 2 esemplari. India: Punjab, Kulu Beas, b. 4000, O. H. Walters, 12.6.1922 (1, BMNH). Pakistan: Phularwan, 11.2.60 / Adult on Pomegranate (1, BMNH). Gruppo di Pachytychius albosparsus (Fàhraeus) Il gruppo presenta la combinazione dei seguenti caratteri: elitre corte, subovali, con squame in massima parte moderatamente allungate, solitamente di due colori ben contra- stanti fra loro, in ogni caso le più scure ricoprenti almeno gran parte delle prime due inter- Revisione dei Pachytychius afrotropicali e orientali 161 strie da dove si dipartono a formare fasce e chiazze più o meno numerose e più o meno lun- ghe; femori posteriori con grosso dente; lobo mediano a forma di corto tubo. Il gruppo è per il momento composto solo da due specie dell'Africa meridionale (distri- buzione in fig. 35) ed appare filogeneticamente correlato con il gruppo di P. haemato- cephalus, da cui differisce essenzialmente, ma nettamente, per la corta forma elitrale. 14. Pachytychius albosparsus (Fàhraeus) (figg. 5, 11, 24, 30) Tychius albosparsus Fahraeus, 1871: 249. Pachytychius albosparsus (Fahraeus), Caldara, 1989: 55. DIAGNOSI. Un Pachytychius del gruppo omonimo con pronoto appiattito soprattutto verso i lati, che sono carenati, con elitre fortemente convesse e con omeri fortemente prominen- ti anteriormente. LOCALITÀ TIPICA. Sud Africa, Caffraria. SERIE TIPICA. Già in precedenza (Caldara, 1989) avevo esaminato in NHMS tre syntypi, designando come lectotypus un d etichettato “Caffraria / I. Vahlb. / Typus / albosparsus Schh.“ e come para- lectotypi due 9 2 etichettate come il lectotypus ad eccezione di “Paratypus”. RIDESCRIZIONE. 3. Lunghezza mm 3,5. Tegumenti neri ad eccezione di apice del rostro, antenne, apice delle tibie e tarsi bruno- rossastri; sul dorso scarsamente visibili fra il fitto rivestimento formato da squame poco allungate (lu/la 3-5), subellittiche, coricate, bianche e nere; le prime sul pronoto formano una fascia longitudinale mediana e due chiazzette laterali basali e due apicali, mentre sulle interstrie elitrali sono raggruppate in numerose chiazze che in parte formano fasce tra- sversali interrotte (dato che tali squame non ricoprono le strie). Strie ben visibili, con sot- tili squame biancastre poco evidenti. Parte ventrale ricoperta da squame bianche e nere (queste ultime ricoprenti gli sterniti III-V e i lati degli sterniti I e II), moderatamente allun- gate (lu/la 4-6), da coricate a subcoricate. Rostro moderatamente più lungo del pronoto (Rlu/Plu 1,17), visto dall'alto lievemente ristretto nel terzo basale, indi a lati paralleli che si allargano un poco in prossimità dell'a- pice, visto di lato fortemente arcuato soprattutto nel terzo basale, pressoché dello stesso calibro dalla base all'apice. Antenne inserite nel terzo apicale; primo segmento del funico- lo lievemente più robusto e circa 1,2 volte più lungo del secondo. Fronte un poco più larga del rostro alla base. Pronoto fortemente trasverso (Pla/Plu 1,52), a lati distintamente carenati e discreta- mente arrotondati alla base e all'apice, più debolmente nella parte centrale, piano ai lati e sul disco nella metà basale, lievemente convesso sul disco nella metà apicale, con punti fitti e profondi in parte riuniti a formare strie soprattutto lateralmente, con intervalli fra i punti stretti e moderatamente lucidi. Lobi oculari debolmente pronunciati. Prosterno debol- mente incavato nel mezzo. Scutello piccolo, scarsamente visibile perché posto su un piano inferiore rispetto a quello del dorso delle elitre. Elitre corte (Elu/Ela 1,17), subovali, a lati arrotondati fin dalla base, con la maggior larghezza un poco oltre la metà, con margine anteriore distintamente concavo, con omeri fortemente prominenti anteriormente, poco più larghe del pronoto (Ela/Pla 1,09), forte- 162 CALDARA mente convesse. Interstrie moderatamente convesse; strie molto evidenti, formate da punti molto profondi, larghe circa la metà della larghezza dell'interstria. Femori posteriori con un forte dente aguzzo; terzo segmento tarsale fortemente bilo- bato e distintamente più largo del secondo. Edeago come in fig. 11. 2. Rostro lievemente più lungo (Rlu/Plu 1,23). Spiculum ventrale e spermateca come in figg. 24 e 30. VARIABILITÀ. La lunghezza è compresa fra 2,7 e 3,7 mm. La variabilità maggiore riguarda il disegno della parte dorsale, dove le squame bianche possono ridursi a formare sul pro- noto solamente una chiazza mediana nella metà basale, mentre sulle elitre possono rico- prire soltanto gli omeri, la parte basale di terza e quinta interstria e formare una fascia tra- sversale mediana che va dalla seconda all’ottava interstria; in casi estremi esse ricoprono solo gli omeri e base della terza interstria. Raramente il pronoto ha lati meno fortemente carenati, più curvilinei e lievemente convessi e la maggior larghezza nel terzo basale. Pla/Plu 1,44-1,54; Elu/Ela 1,14-1,20. Ela/Pla 1,08-1,13. CONSIDERAZIONI E NOTE COMPARATIVE. Il tipo di rivestimento e l’habitus rendono questa specie molto caratteristica e inconfondibile dalle altre specie del genere finora note, com- preso P. tuberculatus l'unica che mostra con essa maggiori affinità morfologiche. NOTE BIOLOGICHE. Non si possiedono dati a riguardo. DISTRIBUZIONE. Sud Africa (Natal). MATERIALE ESAMINATO. Oltre agli esemplari della serie tipica (vedi sopra), altri 8 esemplari. Sud Africa: Natal (1, USNM); Natal, D'Urban [sic] (1, SAMC); Natal, Estcourt (1, SAMC); Umgubada, Sicoast, XI.03 (1, SANC); South Africa, Ntl., Dlinza For. Eshowe, 28.54S 31.28E, 12.1.1984, P. E. Reavell (1, SANC); South Africa, Natal, Vernon Crookes Nat. Res., Umzinto, 30.17S 30.37E, 443 m, 25-26.111.1985, G. L. Prinsloo (1, SANC); con indica- zioni indecifrabili (2, SAMC); senza indicazioni (1, SANC). 15. Pachytychius tuberculatus n. sp. (figg. 6, 12, 23, 29) DIAGNOSI. Un Pachytychius del gruppo albosparsus con protuberanza elitrale a livello della declività posteriore, in corrispondenza dell’apice di quarta, quinta, sesta e settima inter- stria, con pronoto distintamente convesso. LOCALITÀ TIPICA. Sud Africa, Transvaal, Vienna Game Farm Haedspruit. SERIE TIPICA. Holotypus 4 "South Africa, Tvl., Vienna Game Farm Haedspruit, 24.17S 30.58E, 500 m, 17-18.1.1991, R. G. Oberprieler" (SANC). 23 paratypi: 4 dd e 11 £ £ con gli stessi dati del- l'holotypus (9, SANC; 6, RCCM); 1 2 "RSA: E. Transvaal 15 km NE Klaserie, Guernsey Farm, 18- 31.XII.85, S. & J. Peck, woodland, yellow chembaitpantraps" (SANC); 1 4 el 2 "RSA: Transvaal, 15 km NW Klaserie, Guernsey Farm, 19-31.X11.85, S. & J. Peck, woodland evening carnetting" (SANC); 1 2 idem eccetto "Woodland malaise-FIT's" (SANC); 1 9 [Transvaal] "Skukuza NKW. (naby Duba), 15-3-1960, HAD v Schalkwyk" (SANC); 1 9 "Mozambique, F. Muir / Sharp Coll. 1905-313" (BMNH); 2 d d [Mozambico] "Lourenço Marques, F. Muir, 10.VIII.1900 / Sharp Coll. 1905-313" (BMNH). Revisione dei Pachytychius afrotropicali e orientali 163 elongatus leucoloma lyali beardae oberprieleri indicus mungonis = & + + oO # À w viciae Fig. 36. Areale di diffusione del gruppo di P. haematocephalus e località dettagliate di raccolta delle specie afrotropicali e orientali.